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Birds of Danish SPA's

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Contents

1: Designation and management of Danish SPAs

2: Habitats of Danish SPAs

3. Introduction to species accounts

4: Species accounts, part 1 (see below for details)

5: Species accounts, part 2 (see below for details)

6: Discussion

Literature

Appendix 1. Map and list of Danish SPAs

Appendix 2: Table of Reasons for designation

Appendix 3: Table of 1% levels for species occurring in internationally important numbers within Danish SPAs

Appendix 4: Index to species names

Part 1:
Contents of species accounts
Cormorant (Phalacrocorax carbo)
Bittern (Botaurus stellaris)
White Stork (Ciconia ciconia)
Spoonbill (Platalea leucorodia)
Mute Swan (Cygnus olor)
Bewick’s Swan Cygnus (columbianus) bewickii)
Whooper swan (Cygnus cygnus)
Bean Goose (Anser fabalis)
Pink-footed Goose (Anser brachyrhyncus)
Greylag Goose (Anser anser)
Barnacle Goose (Branta leucopsis)
Brent Goose (Branta bernicla)
Common Shelduck (Tadorna tadorna)
Wigeon (Anas penelope)
Common Teal (Anas crecca)
Mallard (Anas platyrhyncos)
Pintail (Anas acuta)
Shoveler (Anas clypeata)
Common Pochard (Aythya ferina)
Tufted Duck (Aythya fuligula)
Greater Scaup (Aythya marila)
Eider (Somateria mollissima)
Long-tailed Duck (Clangula hyemalis)
Common Scoter (Melanitta nigra)
Velvet Scoter (Melanitta fusca)
Goldeneye (Bucephala clangula)
Smew (Mergus albellus)
Red-breasted Merganser (Mergus serrator)
Goosander (Mergus merganser)
Honey buzzard (Pernis apivorus)
Red Kite (Milvus milvus)
White-tailed Eagle (Haliaetus albicilla)
Marsh Harrier (Circus aeruginosus)
Hen Harrier (Circus cyaneus)
Montague’s Harrier (Circus pygargus)

Part 2:
Black Grouse (Tetrao tetrix)
Spotted Crake (Porzana porzana)
Corncrake (Crex crex)
Coot (Fulica atra)
Common Crane (Grus grus)
Oystercatcher (Haematopus oestragalus)
Avocet (Recurvirostra avosetta)
Golden Plover (Pluvialis apricaria)
Grey Plover (Pluvialis squatarola)
Lapwing( Vanellus vanellus)
Red Knot( Calidris canutus)
Sanderling (Calidris alba)
Dunlin (Calidris alpina)
Ruff (Philomachus pugnax)
Bar-tailed Godwit (Limosa lapponica)
Curlew (Numenius arquata)
Spotted Redshank (Tringa erythropus)
Redshank (Tringa totanus)
Greenshank (Tringa nebularia)
Wood sandpiper (Tringa glareola)
Gull-billed Tern (Gelochelidon nilotica)
Sandwich Tern ( Sterna sandvicensis)
Common Tern (Sterna hirundo)
Arctic Tern (Sterna paradisaea)
Little Tern (Sterna albifrons)
Black Tern (Chlidonias niger)
Eagle Owl (Bubo bubo)
Short-eared Owl (Asio flammeus)
Tengmalm’s Owl (Aegolius funereus)
Nightjar (Caprimulgus europaeus)
Kingfisher (Alcedo atthis)
Black Woodpecker (Dryocopus martius)
Woodlark (Lullula arborea)
Tawny Pipit (Anthus campestris)
Bluethroat (Luscinia svecica)
Red-backed Shrike (Lanius collurio)

 

Designation and management of Danish SPAs

Designation

The Council Directive 79/409/EEC of 2 April 1979 on the conservation of wild birds among other bird protection measures obliges (Article 4.1 and 4.2) Member States to classify the most suitable areas for conservation of Appendix I species and regularly occurring migratory species as Special Protection Areas (SPAs).

As a consequence, Denmark in 1983 designated 111 SPAs. The selection of these areas was based on one or both of the following criteria:

  1. The site should be identified as a site of im- portance (mostly breeding sites) for at least one bird species included in Annex I of the Directive.
  2. The site should be identified as a regular staging or wintering area for at least 1% of the flyway population of one or several species not included in Appendix 1.

37 SPAs were designated solely because of their importance for Appendix I species, 15 solely because of high numbers of other staging migrant birds, 59 areas fulfilled both criteria.

One area (no. 79) fulfilled neither1 . For details regarding individual areas see Appendix 2.

General protection of SPAs

In 1994, an executive order2  set up rules for the administration of the Danish SPAs. Besides giving the precise boundaries of the SPAs, the Executive Order inter alia provides rules and guidelines for the administration of the sites by a codification of previous license, planning and management practice and laid down the final boundaries of the Danish SPAs. These guidelines reflect the substantial regulations in the Birds Directive including the amendments added as a consequence of the Council Directive 92/94 of 21 May 1992: the EC Habitats Directive.

No sites were added to or removed from the list; only a number of minor adjustments of existing boundaries were made. In some cases, neighbouring nature sites were included in the SPAs; in other cases, small areas without any significance were excluded. The excluded areas were mainly urban or agricultural lands. Moreover, the Executive Order established that all 27 Danish Ramsar sites in their entirety are included in the SPAs.

The Executive Order states that it is compulsory to consider the SPAs in the regional and municipal planning. No plans will be allowed that bring about deterioration or significant disturbance to the species for which the areas have been designated.

Reserve network

In 1992 78 wildlife reserves was in effect in order to protect breeding, migrating and wintering birds from disturbance and hunting. 48 of these together with a number of other types of reserves totalling 66 reserves were mainly created to protect migrating and wintering waterbirds. The area covered was 705 sq. km. with hunting free zones (525 sq. km. within 32 SPAs). Hunting on the marine territory from motor boat was prohibited in a further 2956 sq. km. of which 1410 sq. km. was within SPAs.

During the revision of the Hunting and Wildlife Management Act in 1992 the Danish Hunters Association, the Danish Ornithological Society and the Danish Society for the Conservation of Nature proposed to establish a network of reserves for staging and wintering waterbirds. Following scientific evaluation of the proposal by the National Environmental Research Institute the National Forest and Nature Agency started implementing the proposal in 1993 with a view to complete the work in 2000. The aim of the network is to provide a better protection against hunting and other recreational disturbances in order to improve the conservation status for waterbirds in Denmark. The original proposal was to establish or enhance 73 reserves in 46 SPAs leading to an increase in hunting and disturbance-free core zones of approximately 650 sq. km. supplemented with an additional 870 sq. km. with regulations of other recreational activities and certain hunting methods.

The new network of reserves within the SPAs is expected to lead to a marked increase in the number of dabbling ducks and geese, due to an increase in the average time spent in Denmark by migrating populations and to population increase due to the improved survival. The reserve network should also lead to improved hunting outside the network. Investigations carried out after the establishment of the reserves support the expectations. At present 53 of the proposed reserves have been established covering 1114 sq. km. in total.

1 This was designated because it holds the only Danish breeding population of Razorbill (Alca torda) and (Common) Guillemot (Uria aalge). The Baltic breeding birds of these species can in fact be considered separate populations (Durinck et al. 1994) and the Danish parts constitute considerably more than 1% of these populations: 13-20% for Guillemot and 4% for Razorbill. However, these populations are mostly resident and thus do not fall under article 4.2 of the Directive.

2 Executive Order No 408 of 25 May, 1994 on the Demarcation and Administration of EC Bird Protection Areas and Ramsar Sites issued by the Ministry of Environment. The executive Order 408 has since been superseded by the Executive Order 782 of 1 November 1998 from the Ministry of Environment and Energy, which integrates the rules for administration of Ramsar Sites, SPAs and proposed Sites of Community Interest (SCIs).

Habitats of Danish SPAs 

The 111 Danish SPAs cover a total area of 976,000 hectares.

The SPAs were selected to cover the full range of important bird habitats in Denmark.

The figures below show the distribution of the total area of the SPAs among the various CORINE land cover classes. The first figure shows the main categories while the following four show the breakdown of the major categories in the relevant sub categories.

As Denmark is internationally most important for staging and wintering waterbirds it is not unexpected that water bodies and wetlands dominate in the total.

 

Arable land makes up the main part of the agricultural areas category but also pastures and various mixed habitats cover considerable areas. The amount of permanent crops is negligible.

 

In the forest and semi-natural areas category various forest types constitutes approximately half the total. Transitional woodland growth accounts for another 7%, some of which might evolve into actual forest while other parts will be cleared to preserve open habitat types. The areas recorded as mixed forest in the CORINE data are rarely mixed stands; rather they consist of a mix of patches of conifers and broad-leaved forest. Most of the forest area is planted and in silvicultural use.

The semi-natural habitats are dominated by natural grasslands and heathlands but also noticeable amounts of dunes and beaches are included.

 

Reflecting the importance of Danish staging areas for migrating waders, the intertidal flats and also saltmarshes dominate the wetland category. The remaining wetlands are peat bogs and inland marshes.

 

 

 

The water bodies category which taken together account for 70% of the total area of the SPAs is dominated by marine areas. Also important are a number of coastal lagoons, some of which are natural while others are the result of partly failed land claims.
The area of freshwater bodies is comparatively small but nevertheless includes very valuable areas.

Introduction to species accounts

Species included here are:

  1. Appendix 1 species which have bred in one or more SPAs 1979-98
  2. All other migrant birds which have occurred in internationally important in one or more SPAs 1979-98.1 

For every species, the following information is included:

Species name
Under this heading the scientific, Danish, and English common names are given.

Red-list and SPEC status DK/Europe/World
DK: According to the Danish red list "Rødliste 1997" (Stoltze and Pihl 1998) and yellow list "Gulliste 1997" (Stoltze 1998).

The 1997 Yellow List is a list of plant and animal species which are decreasing in Denmark, yet still occurring in numbers which are high enough for them not to be included in the 1997 Red List (Stoltze and Pihl 1998), as well as a list of species for which Denmark has a special international responsibility whether they are included in the 1997 Red List or not. Thus, the 1997 Yellow List, covering species which require special attention or are of national responsibility, is closely connected to the Red List. There are three categories in the Yellow List:

  • X (species requiring special attention) is used for declining species not yet rare enough to merit inclusion in the Red List
  • AY are species of national responsibility while breeding
  • AT are species of national responsibility during migration or in winter

Europe: European status (status in brackets means provisional status) and SPEC category according to Tucker and Heath (1994).

The SPEC categories are (from Tucker and Heath 1994):

  • SPEC 1 are species occurring in Europe which are of global conservation concern (threatened, conservation dependent or data deficient)
  • SPEC 2 are species which are concentrated in Europe and have an unfavourable (i.e. Endangered, Vulnerable, Rare, Declining, Localized or Insufficiently Known) conser vation status
  • SPEC 3 are species which are not concentrated in Europe but have an unfavourable (i.e. Endangered, Vulnerable, Rare, Declining, Localized or Insufficiently Known) conservation status
  • SPEC 4 are species which are concentrated in Europe but have a favourable conservat ion status

World: Status according to 1996 IUCN Red List of Threatened Animals (IUCN 1996).

Annexes: The annexes of the Directive on which the species is listed.

General ecology and habitat requirements

Very short description of breeding and/or foraging and roosting habitat.

Occurrence in Denmark

States whether the species occurs as resident, breeding visitor, passage visitor or winter visitor in Denmark.

Trends in Danish population - long-term and 1983-1998

(Breeding birds only).
Development in Danish population this and if possible last century and more detailed for the period since designation of the Danish SPAs. Major threats towards the population(s) also included here.

Trends for relevant populations overall and in Denmark

(Non-breeding birds only).
Overall conservation status for populations utilizing Denmark and development in Danish staging numbers since the designation of the Danish SPAs. Major threats towards the population(s) also included here.

Significance of the Danish population

Size of Danish population in relation to European total. European totals are mainly from Hagemeijer and Blair (1997). The totals given here are the sum of "Europe" and "Russia" as given in Hagemeijer and Blair (1997) excluding Turkey.

Population trend within Danish SPAs and SPA coverage

Included here is a table with data on population within individual SPAs at designation (invariably labelled 1983 in the table though most counts are actually from 1978-1980) and the newest data available from each site. The table is based on data reported to the European Commission in 1983 in conjunction with the designation of the Danish SPAs and data reported to the Commission in 1999 in the NATURA 2000 standard data form. The data reported 1999 are based on a variety of sources, many of which are additional to those in the literature list for this report. The most important additional sources are the many reports from the NERI ecological field stations, the DOF/BirdLife Denmark County reports with results from the 1993-96 bird locality registration project and a wealth of personal comments from a large number of people with first hand knowledge of the various sites.

The column NDY gives the year of the most recent data. Min. and max. BP gives the minimum and maximum for breeding population of the site. NSB is number of staging birds at the site.

The minimum and maximum for breeding pairs both at designation and presently are summed. The resulting totals are to be regarded as rough estimates rather than exact data. As not all areas are counted at the same years pairs changing from site to site may lead to both over and under estimates of total population within SPAs.

Numbers of staging birds are not summed, as the data are not from simultaneous counts.

The coverage (in %) is the fraction of the total Danish population occurring within SPAs. For staging birds the coverage is only given as high (all or nearly all important staging areas are SPAs), intermediate (most important staging areas are SPAs, but some are not) or low (many important sites are not SPAs).

1 A few additional species occurring in internationally important numbers in Denmark are not in included here: Gavia stellata, Gavia arctica, Podiceps grisegena, Uria aalge, Alca torda, Cepphus grylle grylle and Alle alle. These species mainly occur in offshore waters not included in the SPA network.
Likewise not included here are Appendix 1 species only occurring as non-breeding visitors in low densities: Falco peregrinus and Aquila chrysaetos (one pair bred successfully 1999 within SPA no. 7, this is only Danish breeding record). Both these species, though, mainly stage in SPAs in Denmark.
Lastly three rare and irregular breeding Appendix 1 species are not included as data regarding their numbers and distribution are to scanty: Ciconia nigra, Pandion haliaetus and Ficedula parva.

Species accounts, part 1

Cormorant

Species name
Scientific: Phalacrocorax carbo
UK: Cormorant
DK: Skarv

Red-list and SPEC status DK/Europe/World
DK: AY/AT
Europe: Secure, non SPEC
World: Lower Risk (least concern)
Annexes: None

General ecology and habitat requirements
Breeds in trees by lakes or near coast or on ground on predator free islands. Staging birds occur mainly at seacoasts.

Occurrence in Denmark
Two subspecies occur in Denmark ssp. sinensis and ssp. carbo. Ssp. sinensis is mainly a breeding visitor, though some birds stay all year. The number of breeders staying to winter in Denmark is increasing (Laursen et al. 1997). Ssp sinensis also occur as passage visitor, mainly from Sweden. Ssp. carbo occur as passage and winter visitor, probably mainly from Norway (Olsen 1992).

Trends in Danish population - long-term and 1983-1998
The species has bred in Denmark for over 7.000 years (Løppenthin 1967, Grell 1998). The size of the Danish population has for (at least) the last 200-300 years been determined by level of persecution rather than any other ecological factors. In two periods (during much of the 18th century and c. 1870-1938) the species was totally eradicated. Since 1938 the population has increased (slowly at first, dramatically after the species received total protection in 1983) and it reached a maximum of 40.190 nests counted in 1996. In 1997 the first population decrease since 1970 was registered and the number of nests counted fell to 35.829 and an estimated population total of 35,265-37,200 pairs (Eskildsen 1997, Grell 1998).

Current Danish management policy includes that existing colonies are protected, but establishment of new colonies is generally blocked. Illegal persecution by destruction of nests, killing of young and adult birds or release of mammalian predators such as Vulpes vulpes or Mustela vison at formerly predator free islands is probably only threat against existing colonies.

Significance of Danish population
The Danish breeding population equals 24-27 % of total European population.

Population trend within Danish SPAs and SPA coverage and significance of SPAs
The trend within SPAs is identical to the overall sharp increase since 1980 as all main colonies are inside SPAs.

The coverage is thus very high - above 95% - and the management within SPAs is accordingly of the highest importance for the Danish population of the species.

Occurrence in Danish SPAs

Area code Min BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1       326 326   1997  
7   0   3227 3227   1997  
20       1573 1573   1997  
27       66 66   1997 No breeding success
31       2037 2037 4800 1997 Staging numbers are from 1994
36 900 900   4622 4622   1996  
8       120 120 1997   No breeding success
39       1000 1000 2000 1997 No breeding success
43       2905 2905   1997  
53           2320 1992-94  
57           3113 1992-94  
74 P P   3659 3659   1997  
75       12 12   1997 No breeding success
76       2114 2114   1997  
83           2815 1987-89  
85       1212 1212   1997  
87       1833 1833   1997  
88       314 314   1997  
89       3301 3301   1997  
96 1000 1000   2608 2608   1997  
97       1205 1205   1997  
99       1004 1004   1997  
105 0 0   800 800   1997  
108 0 0   394 394   1997  
  1900 1900   34006 34006      

 

Bittern

Species name
Scientific: Botaurus stellaris
UK: Bittern
DK: Rørdrum

Red-list and SPEC status DK/Europe/World
DK: Rare
Europe: (Vulnerable), SPEC 3
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Breeds in extensive reedbeds along lakes and fjords.

Occurrence in Denmark
Resident and breeding visitor, whether the main part of the population is resident or visitor is unknown. Probably also occurs as passage or as winter visitor from the Swedish population.

Trends in Danish population - long-term and 1983-1998
After a period of expansion of breeding area and population growth in the beginning of the century, the Danish population suffered severe setbacks from 1930-1950 mainly caused by drainage of several important breeding localities. The population remained low (10-20 "pairs") until the beginning of the 1970s. During the years 1978-81, the population was estimated at 24-40 "pairs". From the 1980s, the Danish population of the species has increased rapidly and the present (1993-96) population estimate is 150-200 "pairs". The reasons for this dramatic recovery are not clear. Improved water level management at the key site Vejlerne (SPAs no. 13 and 20), which harbour roughly half the Danish population and the total protection of Grey Heron (Ardea cinerea) since 1981, thereby limiting accidental killing as a result of misidentification, are probably contributing factors.

Overlaying the long-term trend are year-to-year fluctuations reflecting increased mortality during harsh winters.

Significance of the Danish population
The Danish population is only a small part (probably less than 1%) of the total European population which was recently estimated at 20,044-41,669 "pairs" (Hagemeijer and Blair 1997). Nevertheless, the situation in Denmark is of considerable interest as it is one of the very few European countries with a positive population trend.

Population trend within Danish SPAs and SPA coverage
The larger part of the Danish population is concentrated within a few areas, all of which are SPAs. The recent substantial increase has mainly taken part within the key sites, though some expansion to new localities has also occurred. The population increase in the most important area Vejlerne (SPAs no. 13 and 20), harbouring nearly half the total Danish population, is partly due to management changes with more favourable water levels. At the only site showing a marked decrease Vidåen, Tøndermarsken and Saltvandssøen (SPA no. 60) the opposite is true, as current lower water levels are less favourable for the species than formerly.

At designation the population was covered almost fully (>90%). With the recent expansion to new breeding areas this coverage has fallen and is presently somewhere in the range 70-90%.

The SPAs still harbour the main part of the Danish population and appropriate management here is of very high importance for the favourable conservation status of the species in Denmark. The larger relative increase outside than inside SPAs probably reflects the spread to more marginal habitats of a rapidly increasing population and the effect of small restoration projects rather than any general management problems within SPAs.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
13 9 9   40 40   1995  
20 9 9   20 20   1995  
21       1 1   1987-89  
29       1 1   1995  
37       1 1   1993-94  
38       1 1   1996  
41       3 3   1995  
43 4 8   5 9   1993-94  
47       0 0   ? No breeding since 1988
53       3 4   1993-94  
56 0 0   1 1   1993-94  
58   1   0 1   1989  
60 11 16   3 3   1996  
63 1 1   0 1   1989  
65 1 1   2 3   1994  
71 1 1   0 0   1996  
72 1 1   1 1   1995  
80       0 2   1992  
83       1 2   1994  
87       35 35   1995  
89       2 2   1996  
100       1 1   1994  
106 1 1   1 1   1993  
Total 38 48   122 133      

 

White stork

Species name
Scientific: Ciconia ciconia
UK: White Stork
DK: Hvid stork

Red-list and SPEC status DK/Europe/World
DK: Endangered
Europe: Vulnerable, SPEC 2
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
White storks nest on buildings and forages in wetland and natural grasslands.

Threats
Main threats are. Also collision with power lines and hunting or poisoning during migration and in winter quarters are important causes of mortality (Tucker and Heath 1994, Grell 1998).

Occurrence in Denmark
The species is a breeding visitor to Denmark.

Trends in Danish population - long-term and 1983-1998
It is estimated that the Danish population numbered 10,000 pairs 200 years ago.

Mainly because of inappropriate water level management (i.e. decreased spring floodings) and general agricultural intensification in river valleys availability of main prey (amphibians) have decreased. This has resulted in an almost steady decline of the Danish population. The decline started mid 19th century and around the year 1900 only 4-500 pairs remained. A short-lived, partial recovery occurred during the 1930s resulting in a population of 1,100 in 1939. Since then, the story is one of continual decline to the 1997 status of 4 pairs (Grell 1998). Only two pairs bred 1999.

Some ongoing restoration efforts, especially the Skjern Å valley project, might provide better opportunities for the species in the future.

Significance of the Danish population
The remnant and possibly doomed Danish population is only of marginal significance in an international context. The European population was recently estimated at 104,154-111,710 pairs (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
None of the 3 remaining pairs (1998) have their nest inside SPAs, but 2 are just on the border of SPAs and are almost totally dependent on the SPAs (no. 13 and 51 respectively) as foraging areas. The third pair nests approximately 3.5 km south of SPA no. 1 and partly forages within this area.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
13           5 1996  
20           5 1996  
44           1 1996  
60 1 1   1 1   1996 Not breeding 1997-98
82           4 1993  
Total 1 1   1 1      

 

Spoonbill

Species name
Scientific: Platalea leucorodia
UK: Spoonbill
DK: Skestork

Red-list and SPEC status DK/Europe/World
DK: Extinct*
Europe: Endangered, SPEC 2
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Nests on islands free of ground predators, dense reed beds and willow thickets. The species feeds on a wide variety of aquatic prey and even some plant material (Snow and Perrins 1998). As the species is a tactile feeder high abundance of prey is required. Foraging takes place up to 25 km from nest site (Tucker and Heath 1994), though breeding success decreases with distance (Grell 1998).

Occurrence in Denmark
The species is a scarce and irregular breeding visitor.

Trends in Danish population - long-term and 1983-1998
The species has bred in Denmark in 1900, 1919 (2-4 pairs), 1928-29, 1942-43 (4-5 pairs), 1962-69 (1-7 pairs) and 1996-1999 (Løppenthin 1967, Olsen 1992 and Grell 1998).

Significance of the Danish population
The Danish population is a marginal extension of the increasing Dutch population (833 pairs in 1996, Snow and Perrins 1998). Breeding failure as a result of low water levels in 1996 at the important Oostervaardersplassen colony resulted in flocks of displaced birds turning up all over northern Europe and re-colonization of Denmark (Grell 1998). The total European population is estimated at 4,931-5,984 pairs (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
The Danish population is wholly dependent on SPAs both for nest sites, foraging and as post-breeding staging area.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1       3 3   1997  
13       1 1 13 1996 Only bred 1996 not since
Staging number 1998 data
20           4   1997
Total       3 3      

 

Mute Swan

Species name
Scientific: Cygnus olor
UK: Mute Swan
DK: Knopsvane

Red-list and SPEC status DK/Europe/World
DK: Yellowlisted as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Breeds mostly at freshwater sites with well developed submerse vegetation. Also, sometimes colonially, at brackish fjords. Birds staging during moult and in winter occur mostly in fjords and other shallow waters with beds of eelgrass (Zostera).

Occurrence in Denmark
The species is a breeding resident and a passage and winter visitor (Olsen 1992). Denmark is most important as moulting (July and August) and wintering area for a large part of the Baltic population.

Trends for relevant population(s) overall and in Denmark
There are no information regarding trends in Danish staging population during moult or in winter. In the period 1987-91, the number staging in Denmark during moult was estimated at 34,000-59,000 individuals, with 37,000-73,0000 individuals wintering (Jensen 1993). The total northwest European population is increasing (Scott and Rose 1996).
The breeding Danish population is estimated at 5,000 pairs and is believed to be stable (Grell 1998).

Significance of the Danish population
During moult up to 28% and in winter up to 35% of the total northwest European population of 210,000 birds (Scott and Rose 1996) stage concurrently in Denmark.

Population trend within Danish SPAs and SPA coverage
Numbers of staging birds within individual SPAs are fluctuating but total staging population within SPAs seems reasonably stable. The coverage is high as most but not all areas of high densities (Durinck et al. 1994) are covered by the SPA network.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1     1500     2939 1993  
15     5000     1202 1993-94  
31     2000     428 1993-94  
36     400         No new data
38     2000     55 1996  
43 900 900 4300 160 170 1000 1993-94  
71     6000     5860 1989  
72     1       1983  
75 175 175 10000     3700 1993-94  
76     2000     120 1993-94  
81     3000     5900 1991  
83     5000     13097 1994 Data from part of area only
84     8000     4200 1991  
85 230 230 6000     9757 1993-94  
86     3000     870 1995 Data from part of area only
88     8000     3730 1994  
89     6000     8675 1991  
95     3000     228 1987-89  
96     3000     2440 1987-89  
105 300 300 3500-3700   893 3062 1998 Breeding data are from 1993
110     2300     2403 1994  
111     1600     1114 1990-91  
                 
                 

 

Bewick’s Swan

Species name
Scientific: Cygnus (columbianus) bewickii
UK: BewickŽs Swan
DK: Pibesvane

Red-list and SPEC status DK/Europe/World
DK: Yellow-listed as AT
Europe: Localized, SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Formerly staging birds mainly foraged on submerse vegetation in shallow lakes and coastal waters. The reduction or disappearance of submerse vegetation (Characeans, Zostera and Potamogeton) at some sites as a result of eutrophication has, though , led to increased instance of foraging in autumn sown cereals (Grell 1998).

Occurrence in Denmark
The species is a passage and to a lesser extent a winter visitor to Denmark (Grell 1998).

Trends for relevant population(s) overall and in Denmark
The size of population breeding in western Siberia and wintering in northwest Europe is estimated to be 17,000-25,000 birds. The population is thought to be increasing (Scott and Rose 1996).

In Denmark 4,000-6,000 birds occur as passage visitors in autumn and up 1,000 winter in mild winters (Grell 1998).

Significance of the Danish population
The 4,000-6,000 birds occurring at peak of autumn passage equals16-35% of European total.

Population trend within Danish SPAs and SPA coverage
Staging numbers fluctuates markedly as counting timing is not always optimal for getting peak numbers. Overall, the numbers in SPAs are probably roughly stable. The coverage is intermediate; most but not all important areas as shown in Laubek 1995) are SPAs. Notable exceptions are Bolle and Try Enge, Store Vildmose and Øland-Attrup. The management of the SPAs is despite these gaps of high importance for the Danish staging population.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1   4086 1995  
8 230   1994  
13 318 377 1994  
19 600 316 1994  
20 500 403 1993-95  
21 100     No new data
23 600 140 1993-94  
24 130 723 1993-95  
38 1500 101 1996  
39   105 1993-96  
41 1000 1000 1995  
43 1100 900 1993-94  
56 500 250 1993-94  
60 225 332 1992-94  
97 100 5 1994  
100 400 76 1987-89  
101   76 1987-89  

 

Whooper Swan

Species name
Scientific: Cygnus cygnus
UK: Whooper Swan
DK: Sangsvane

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, SPEC 4 (winter)
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Staging Whooper Swans forage in winter cereals or oilseed rape, fields with waste grain potatoes or beets, on flooded meadows and in waterbodies with abundant bottom macrophytes.

Occurrence in Denmark
The species is a winter visitor to Denmark.

Trends for relevant population(s) overall and in Denmark
The northwest continental Europe winter population is increasing rapidly and is presently estimated at 40,000 birds (Scott and Rose 1996). The Danish winter population is estimated at nearly 16,000 birds (Grell 1998) and has likewise increased in recent years.

Significance of the Danish population
16,000 birds equal 40% of total northwest continental European population.

Population trend within Danish SPAs and SPA coverage
The numbers counted in individual SPAs fluctuate markedly but the total staging numbers in all SPAs is probably roughly stable. The coverage is intermediate, not all major staging sites are SPAs (see Laubek 1995). As a result of the recent shift in foraging habitat some agricultural areas have become of importance for the species. These areas, which include Bolle and Try Enge, Store Vildmose, Søby Sø and surrounding areas and the reclaimed areas of Lammefjorden are not SPAs. Nevertheless is appropriate management of the SPAs of high significance for the Danish staging population.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 1600 4320 1995  
7   528 1995  
8 700 9 1994  
10   200 1987-91  
12 900 238 1994-96  
13 400 885 1994  
15 2000 2000 1995  
19 360 361 1994  
20 300 403 1993-95  
21 400 143 1987-89  
22   430 1993  
23 700 414 1996  
24 400 885 1995  
30   260 1995  
31 200 76 1995  
33   128 1995  
35   80150 1993-95  
37   100-200 1997?  
38 1000 119 1994  
39 200 76 1996  
41 500 367 1996  
43 900 100 1993-94  
44   196 1996  
47 1000 36 1995 Only small part of area counted
56 300 471 1994  
60 375 941 1992-94  
71 600 600 1996  
72 1   1983  
75 300 300 1993-94  
76   250 1993-94  
81 900 607 1987-89  
82   106 1995  
83 300      
84 2000 175 1993-94  
85 700 400 1994  
86 350 34 1993-95 Only part of area counted
88 700 424 1996  
89 1800 270 1987-89  
91 330 106 1995  
94   266 1996  
95 700 30 1987-89  
96 200 156 1987-89  
97   132 1996  
99 150 341 1994  
100 190 1993 1994  
105 250-600 462 1998  
110 240 1 1994  

 

Taiga Bean Goose

Species name
Scientific: Anser fabalis fabalis
UK: Taiga Bean Goose
DK: Skovsædgås

Red-list and SPEC status DK/Europe/World
DK: Yellow-listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1

General ecology and habitat requirements
In southeastern Denmark staging Taiga Bean Geese mainly forage on winter cereals, seed grass and waste sugar beet, while the small group occurring in northern Jutland utilises pastures, stubble fields and winter cereals (Madsen et al. 1999).

Occurrence in Denmark
Occur as a passage and (mainly) winter visitor.

Trends for relevant population(s) overall and in Denmark
The total population of the subspecies has been increasing at least since 1960. In the main staging sites in southern Sweden the numbers have increased from 12,800-16,800 in the late 1950s to 60,000-80, 000 in recent years. The total population is now estimated at 90,000-110,000 individuals and has apparently stabilised (Madsen et al. 1999).

Two sub-populations occur in Denmark. A smaller one on in northern Jutland occurs mainly during passage, some of these birds winter in England. Higher numbers occur in southeastern Denmark. These birds are mainly winter visitors. Highest number during cold winters or spells where high numbers are displaced here from Scania (Sweden) (Jørgensen et al. 1994).

Significance of the Danish population
In mild winters 8,000-10,000 birds winter in southeastern Denmark, while 18,000-32,000 (or up to 35% of total population) occur in cold winters (Madsen et al. 1999).

Population trend within Danish SPAs and SPA coverage
As stated by Madsen et al. (1999) for the total Danish population no trends within the SPAs are discernible apart from large yearly fluctuations at the Southeast Danish sites in relation to weather conditions.

Nearly all main roosting sites are SPAs, but foraging to a large extent takes place outside SPAs mainly in cultivated areas. Hence, coverage for roosting sites is high while it is low or intermediate for foraging areas.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
13 900 477 1994  
16 1200 290 1988  
20   999 1993-95  
22 800 1050 1993-94  
65 1200      
81 600-900 600 1994 Only part of area counted
82 2000 2000 1993-94  
83 3000      
91 2200 2560 1994  
93 600-1200 2210 1993-94  
96 1200 1700 1987-89  
110 520      

 

Pink-footed goose

Species name
Scientific: Anser brachyrhyncus
UK: Pink-footed Goose
DK: Kortnæbbet gås

Red-list and SPEC status DK/Europe/World
DK: Yellow-listed as AT
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Pink-footed Geese in autumn feed mainly on stubble fields and to a lesser extent on winter cereals. Later they switch to pastures and during frost periods to winter cereals. In spring some newly sown areas of spring cereals and peas are also utilised. Earlier saltmarshes and semi-natural grasslands were also utilised, but during the 1990s the geese seem to have abandoned these habitats (Madsen et al. 1999).

Occurrence in Denmark
Occur as passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The birds occurring in Denmark come from the population breeding in Svalbard, which has been increasing since the 1950s. It is estimated that the population totalled 10,000-12,000 birds in the 1950s while it presently numbers 32,000-37,000 birds (Madsen et al. 1999).

Significance of Danish population
At peak time from late February to late April virtually the whole population is concentrated in Denmark. In spring 14 major sites can be identified, while in autumn the geese in recent years concentrate at only two sites (Madsen et al. 1999) in response to hunting pressure. The present level of hunting has not impeded the growth of the population during 1980s and 1990s (Madsen et al. 1996).

Population trend within Danish SPAs and SPA coverage
As the total population occurs in Denmark at peak time and all major roosts are SPAs the trend for total population, Danish population and population within SPAs are virtually identical. Some foraging, though, takes place outside areas. Nevertheless, coverage is high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 450 7    
7   420 1993  
13 123 2660 1994  
17 150      
20   3613 1993-95  
22 1500 350 1993-94  
23 3000 309 1993-94  
38 3750 5000 1996  
39 3500 4500 1993-96  
41 15000 11100 1995  
43 15000 8000 1993-94  
51 500 1000 1993-94  
56 28500 30000 1993-94  
57 7000 10000 1992-94  
60 3500 430 1992-94  
65 1300      
67 9000 18500 1994-95  

 

Greylag goose

Species name
Scientific: Anser anser
UK: Greylag Goose
DK: Grågås

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1

General ecology and habitat requirements
Staging Greylags during August-September forage on waste grain in stubble fields, grass, unharvested peas and waste potatoes. In October the birds in southeast Danish sites mainly forage on waste sugar beets and locally on winter rape and cereals (Madsen et al. 1999).

Occurrence in Denmark
The species occurs in Denmark as breeding visitor, a passage visitor and also (at least in mild winters) in lower numbers as winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European population has increased dramatically from around 30,000 birds in the mid 1970s to probably more than 200,000 at present. The most important factors influencing survival and condition are amount of hunting and ecological conditions (especially amount of rainfall) in Spanish winterquarters. The number of birds staging in Denmark at peak time in autumn has likewise increased from around 25,000 birds in 1981 to more than 40,000 birds in the 1990s (Madsen et al. 1999). In 1996, the count went as high as 60,000 (Pihl et al. 1997). As the number of Greylags at the staging sites have only increased moderately in southeastern Denmark and even decreased slightly in the central part of the country the overall large increase in numbers is mostly a result of a rapid increase in the number of birds staging in western Jutland. These birds are mostly from the population breeding in Norway (Jørgensen et al. 1994).

The number of birds breeding in Denmark was recently estimated at 3,500-4,000 pairs or possibly more (Grell 1998).

Significance of the Danish population
Denmark is very important for staging Greylags, mainly in autumn.

20-30% of the northwest European population can occur simultaneously in Denmark.

A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
Most of the major staging areas are SPAs, hence the coverage is high and trends within SPAs are almost identical to overall national trends. Foraging, though, to a large extent takes place outside areas. In some instances (notably SPA nos.82 and 101) the SPAs only cover parts of larger continuous staging sites. Nevertheless, adequate management of SPAs is of vital importance for the Danish population of the species.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Com-
ments
1     1000     650    
13 700 800 1000 800 850 2064 1994 Breeding data include area 20
20     400     1978 1993-95  
21     450          
22           1700 1993-94  
38           4000 1996  
56           22000 1993-94  
57           10000 1992-94  
74     4000     700 1993-94  
78     3700     720 1996  
82     800     2000 1993-94  
83           4000 1987-89 Large numbers not regular
87 280 280 4000     6675 1995  
91     600     530 1995  
95     500     1495 1991  
96     3000     11700 1987-89  
99     3000     1500 1994  
100 48 58 3000     2024 1994  
101 1 1 2000     1540 1987-89 Up to 4.365 if adjacent areas included
105     5000     1082 1997  
110     630     6365 1994  

 

Barnacle Goose

Species name
Scientific: Branta leucopsis
UK: Barnacle Goose
DK: Bramgås

Red-list and SPEC status DK/Europe/World
DK: Rare, i (breeding)
Europe: Secure (breeding), Localized (winter), SPEC 4 (breeding), SPEC 2 (winter)
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Staging Barnacle Geese feed on coastal saltmarshes and inland pastures, in recent years increasingly also in winter cereals (Madsen et al. 1999). The newly established breeding population occur on islands free of mammal predators.

Occurrence in Denmark
The species is mainly a passage and in mild winters also a winter visitor to Denmark. Recently the rapidly expanding population breeding on islands and islets in the Baltic Sea has reached Denmark.

Trends for relevant population(s) overall and in Denmark
The birds occurring in Denmark belong to the Russian and Baltic population. This population has increased 25-fold since the 1950s, where the population experienced an all-time low of around 10,000 birds. The mid-winter count of January 1997 gave a result of 267,000 birds. The small Baltic breeding segment has experienced a particularly rapid growth since its foundation in 1971 to presently around 13,000 birds (Madsen et al. 1999).

There are no serious threats towards the staging visitors in Denmark. The small newly established breeding population is vulnerable to disturbance and especially to eventual invasion of mammalian predators (e.g. Fox (Vulpes vulpes) or American Mink (Mustela vison)) on formerly predator free islands.

On flyway level the most important threats are believed to be large scale development of oil and gas industry at the Russian breeding grounds (Madsen et al. 1999).

Significance of the Danish population
Up 20,000 birds or 7,5% of flyway total winter in Denmark in recent years, probably as a result of a string of mild winters (Madsen et al. 1999). Even higher numbers occur during migration periods.

Population trend within Danish SPAs and SPA coverage
Only three sites regularly support numbers of international importance (Madsen et al. 1999) and all are SPAs (nos. 60, 67 and 89). Also most areas occasionally holding large numbers are within the SPA network. The coverage is thus high.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Com-
ments
15           800 1993-94  
23           1160 1993-94  
31           732 1995  
57     3000     7700 1992-94  
60     1150     30000 1997  
67           19000 1994-95  
75           540 1993-94  
76           114 1995  
81           343 1995  
82           3500 1993-94  
89           6000 1996  
96           1215 1987-89  
110     2000 50 50 1523 1999 Staging numbers are from 1994

 

Brent Goose

Species name
Scientific: Branta bernicla
UK: Brent Goose
DK: Knortegås

Red-list and SPEC status DK/Europe/World
DK: AT
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Brent Geese forages on algae, Zostera beds (most important in autumn), saltmarsh vegetation and recently also on farmland (Tucker and Heath 1994, Madsen et al. 1999)).

Occurrence in Denmark
Occur as passage and winter visitor. Two different populations/subspecies are involved namely Dark-bellied Brent Goose (ssp. bernicla) breeding in Siberia and the Svalbard breeding population of Light-bellied Brent Goose (ssp. hrota).

Trends in Danish population - long-term and 1983-1998
Both populations occurring in Denmark have as a result of total protection since 1972 increased. The population of Dark-bellied Brent has increased from 40,000 individuals in the 1960s to 200,000-300,000 in the 1990s. The growth rate of this population has decreased recently, and the population might now be stabilizing, though this is still uncertain (Madsen et al. 1999).

For the Svalbard population of Light-bellied Brent Geese, the increase has been less dramatic: from 2,000 individual in the 1960s to 4,000-5,800 in the early 1990s (Madsen et al. 1999). Up to nearly 4,000 birds have been recorded simultaneously in Denmark (Jørgensen et al. 1994).

The number of autumn staging Dark-bellied Brent Geese in Denmark is not well known, but during 1980-82 10,000-40,000 individuals were registered during nation-wide counts in October. Clausen et al. (1997) gives the maximum number of simultaneously staging birds as 47,000. In mild winters 2,000-4,000 stay within the country and during the spring peak period an estimated 18,000-24,000 birds stage. The number of this subspecies staging in Denmark has not increased proportionately with increase of the total population (Jørgensen et al. 1994).

Significance of the Danish population
Denmark is of the highest importance for the conservation of the small Svalbard population of Light-bellied Brent Goose. 50-75% of the total population winter in Denmark in mild winters. At least 80% of the population is in Denmark during spring peak.

47,000 Dark-bellied Brents equal 16–23% of total population. A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
All important staging areas for Light-bellied Brent Goose are SPAs and the SPAs thus at times holds at least 75% of the total population.

The most important Danish staging areas for Dark-bellied Brent Goose are SPAs, but many regular sites of secondary importance exist outside the SPA-network, still the coverage is high.

The SPA network is thus of principal importance for the conservation of both populations staging in Denmark especially the Svalbard population of Light-bellied Brent Goose.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1   1200 1994 ssp. hrota
2   900 1993-94 ssp. hrota
3 1200 1600 1987-91 ssp. bernicla
15 1900 3500 1993-94 mainly ssp. hrota, but figure includes a smaller number of ssp. bernicla
23   512 1996 ssp. hrota
26 300 595 1991 ssp. bernicla, some years also small numbers of ssp. hrota
27 1000 2000 1995 ssp. hrota
38 2300 850 1996 ssp. hrota
39   1300 1993-95 ssp, hrota, only parts of area counted
40   380 1995 ssp. hrota
52 17000 10000 1993-94 ssp. bernicla
52   2000 1993-94 ssp. hrota
53   6500 1992-94 ssp. bernicla
53   1598 1992-94 ssp. hrota
57   2000 1992-94 ssp. hrota
57 25000 13300 1992-94 ssp. bernicla
60 3500 6811 1992-94 both subspecies
65 6000 1400 1991-95 mainly ssp. bernicla
67   5050 1988-89 mainly ssp. bernicla
71 2000 4000 1987-89 ssp. bernicla
83 2000 214 1994 ssp. bernicla

Common Shelduck

Species name
Scientific: Tadorna tadorna
UK: Common Shelduck
DK: Gravand

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: None

General ecology and habitat requirements
Staging Shelducks occur at coastal mudflats, sheltered seacoasts and in coastal lagoons and estuaries.

Occurrence in Denmark
Occur as breeding, moulting and wintering visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European population increased by at least 50% between the late 1960s and the late 1980s. However, in recent years, the rate of increase has decreased, and the population might now be stabilizing. It is presently estimated at 300,000 birds (Scott and Rose 1996). The Danish breeding population was recently estimated at 1,650-2,650 pairs (Grell 1998).

Significance of the Danish population
Totally up to 45,000 birds can occur simultaneously in Denmark (Clausen et al. 1997), this equals 15% of flyway population. The Danish winter population probably increased between 1968-73 and 1987-89 (Laursen et al. 1997).

The Dutch-German-Danish Waddensea is the main moulting and wintering area for the northwest European population. During moult most birds are concentrated in the German part of the area. The relative importance of the Danish Waddensea increases in the post moulting period (autumn) where more than 20,000 birds normally occur, during mild winters similar numbers are present (Grell 1998). Outside the Waddensea region internationally important numbers only occur in one area in Kattegat.

Population trend within Danish SPAs and SPA coverage
At the keysites in the Waddensea (SPAs nos. 52,53,57 and 60) and Kattegat (no. 15) the numbers seem to have been stable or increasing, while staging numbers at some sites of secondary importance have possibly decreased since designation. All Danish sites important for the species are included in the SPA network. Coverage thus high

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
1 1000    
2 1000    
8 1500 195 1994
12 1500 212 1994-96
13 1000 343 1994
15 4000 5670 1993-94
23 2200 1493 1993-95
24 1400 271 1993-95
43 2000 400 1993-94
52 3000 2000 1993-94
53   7600 1992-94
57 35000 35584 1995
60   6466 1992-94

 

Eurasian Wigeon

Species name
Scientific: Anas penelope
UK: Eurasian Wigeon
DK: Pibeand

Red-list and SPEC status DK/Europe/World
DK: Vulnerable (breeding) and Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Wigeons forage mainly in saltmarshes, intertidally in eelgrass beds and in recent years also in winter cereals.

Occurrence in Denmark
The species is a rare breeding visitor and abundant passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
In contrast to the development in the other West Palaearctic populations the northwest European population has apparently increased steadily for the last 20 years and is presently estimated at 1,250,000 birds (Scott and Rose 1996). Concurrently the estimates of numbers staging simultaneously in Denmark have risen from 45,000 in 1987-91 (Jensen 1993) to 75,000 in recent years (Grell 1998).

The present level of hunting has not impeded a large growth in the population during 1980s and 1990s. Despite this, the species is regarded as susceptible to disturbance as a result of hunting as it is gregarious and mainly occur by the coast (Madsen et al. 1996).

The positive development in Denmark is probably a combined result of the general development in the population and the creation of many new hunting and disturbance-free core zones within the SPAs.

Significance of the Danish population
Up to 6% of total population stage concurrently in Denmark.

A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
The trend within SPAs is similar to the overall trend, at some sites where new hunting and disturbance-free core zones have been established, though, the increase in numbers has been much higher than average.

All main staging areas are SPAs; the coverage is thus high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
1 16000 26650 1993
8 2000    
12 8000 2860 1994-96
13 6000 4450 1994
23 11500 3548 1996
36 P 592  
38 8000 981 1996
39   825 1994
43 30000 9500 1993-94
52   8000 1993-94
53 10000 2305 1992-94
55 8000 1726 1996
57 54000 51178 1995
60 16800 5962 1992-94
65 28000 1950 1991-95
67   10920 1988-89
89   35000 1996
99 3000    
110 8200 3245 1994

 

Common Teal

Species name
Scientific: Anas crecca
UK: Common Teal
DK: Krikand

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Common Teals occur at both freshwater and brackish wetlands.

Occurrence in Denmark
The species is an uncommon breeding visitor and an abundant passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European winter population has increased slowly since 1967 and is presently estimated at 400,000 individuals (Scott and Rose 1996).

As the species is very gregarious and mainly occurs, near-shore it is particularly sensitive to disturbance from hunting (Madsen et al. 1996). Accordingly it is one of the species benefited by the creation of the hunting and disturbance free core zones (Madsen et al. 1995).

Significance of the Danish population
Up to 50,000 birds (or 12,5% of population total) stage simultaneously in Denmark (Grell 1998).

A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
Numbers counted at Danish SPA sites seems to have declined in recent years at several sites. Some of this decrease (e.g. areas 38 and 43) is probably real and result from decreased habitat quality as a result of eutrophication. Some of the apparent decrease, though, might just be random fluctuations. At other sites, numbers have increased as a result of creation of new hunting and disturbance-free core zones.

All major staging sites are SPAs, the coverage is thus high and appropriate management of SPAs is therefore of high importance for the Danish staging population.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
01 5000 2378 1993  
05     1994  
13 7000 3642 1994  
14 9000 0 1993-94  
17     1994  
22     1994  
23 3000 956 1996  
29 2000 150 1994  
37     1995  
38 5000 2770 1996  
41 5000 640 1996  
43 11000 6900 1993-94  
44     1993  
57 17000 3000 1992-94  
65 11000 200 1991-95  
67   4165 1988-89  
83   4900 1993  
91 2000     No new data
99 5500 450 1994  
100 2000 103 1987-89  

 

Mallard

Species name
Scientific: Anas platyrhyncos
UK: Mallard
DK: Gråand

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1,III/1

General ecology and habitat requirements
Staging Mallards occur in almost all wetland types apart from off shore. Highest numbers occur at freshwater sites and at sheltered coastal sites.

Occurrence in Denmark
The species occurs as resident, breeding, passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European population is thought to be stable at around 5,000,000 individuals (Scott and Rose 1996).

Significance of the Danish population
As the total population is very large, no Danish sites reach 1% levels. At a few sites, however, the species alone satisfies the 20,000 waterbirds criterion.

Up to 183,000 birds stage simultaneously in Denmark and a considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
All sites with high numbers of staging birds are SPAs. As the species is ubiquitous, total numbers outside SPAs are also considerable. The coverage is thus intermediate.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
43 22000     No new data
57 38000     No new data

 

(Northern) Pintail

Species name
Scientific: Anas acuta
UK: (Northern) Pintail
DK: Spidsand

Red-list and SPEC status DK/Europe/World
DK: Vulnerable (breeding) and Yellow listed as AT
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Pintails occur mainly in coastal areas such as sheltered fjords, coastal lagoons, estuaries and saltmarshes.

Occurrence in Denmark
The species is a scarce breeding visitor and a more abundant passage and (in mild winters) winter visitor (Olsen 1992).

Trends for relevant population(s) overall and in Denmark
The northwest European wintering "population" (though hardly discrete from the rest of the West Palaearctic population) is decreasing and is presently estimated at 60,000 birds, while the total West Palaearctic population is thought to number close to 2,000,000 birds (Scott and Rose 1996).

Main threats against the European population are believed to be habitat destruction and excessive hunting (Tucker and Heath 1994).

The overall effect of hunting on the population status is unknown, but as the species is gregarious and mainly occur near coast, it is particularly susceptible to disturbance as a result of hunting (Madsen et al. 1996). Accordingly it is one of the species most benefited by the creation of the hunting and disturbance free core zones (Madsen et al. 1995).

Significance of the Danish population
In the period 1987-89 up to 8,600 birds (14% of northwest European total) staged simultaneously and up to 6,000 (10% of northwest European total) occurred in winter (Jensen 1993). The Danish winter population is believed to have increased as counts from 1968-73 are much lower than 1987-89 (Laursen et al. 1997). A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
The staging numbers within Danish SPAs seem to have decreased in recent years. The reasons are unknown, but some of the observed decreases are probably part of the general decline while some of the observed changes might be random fluctuations. In some SPAs (e.g. no. 43) decreased habitat quality as a result of eutrophication of waterbodies is probably at least partly responsible for the observed declines.

All main staging areas are SPAs and coverage is thus high. Appropriate management of SPAs is of high importance for the Danish staging population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Com-
ments
1 1 1 1100     12 1993  
13 13 18            
19 1 1            
23 6 6 1800     870 1996  
30 1 1            
38 1 1 2000     289 1996  
41 1 1 11000     1120 1994  
43 10 13 18000     2000 1993-94  
47 10 10 1000          
53           2960 1992-94  
56     5000     3000 1993-94  
57 10 10 8000     4412 1995  
60     1300     900 1991-95  
65     5000     200 1991-95  
71 24 24   25 30   1996  
76 1 1   3 3   1994  
81 1 1   15 22   1996-98  
82 1 1   0 0   1993  
83 10 15   8 8   1993-96  
84 1 1   4 4   1993-94  
85 1 1   1 1   1996-98  
88 8 8   0 0   1993-94  
89 20 20   13 14   1996-98  
95 1 1   0 1   1988  
96 12 12 1500 4 5 785 1987-89  
99 1 1           No new data
100 1 1   0 0   1993-94  
110 5 5   3 3   1993  
111 1 2   0 0   1993  

 

(Northern) Shoveler

Species name
Scientific: Anas clypeata
UK: (Northern) Shoveler
DK: Skeand

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Shovelers occur at coastal lagoons, fjords and freshwater lakes.

Occurrence in Denmark
Occur as breeding, passage and (in low numbers) winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European wintering population appears to have been relatively stable during the period 1982-93 and is estimated to number 40,000 birds (Scott and Rose 1996). The Danish breeding population was recently estimated at 800-1,000 pairs (Grell 1998).

The overall effect of hunting on population size is unknown, but as the species is gregarious and coastal, it is susceptible to disturbance from hunting (Madsen et al. 1996). Accordingly it is one of the species most benefited by the creation of the hunting and disturbance free core zones (Madsen et al. 1995).

Significance of the Danish population
Only a few percent of total northwest European population can occur simultaneously in Denmark (Grell 1998). A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

Population trend within Danish SPAs and SPA coverage
The trend within SPAs is variable. At some sites staging numbers have decreased at other increased. Some of the observed changes are probably just random fluctuations. Some of the decreases might be caused, though, by decreased habitat quality (e.g. at SPA no. 43). The large increase at SPA 89, from 100 birds in 1987-88 to 1,700 in 1993 is undoubtedly a result of the new hunting and disturbance-free core zone created here (Madsen et al. 1995).

All major staging sites are SPAs and the coverage is thus high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 250     No new data
13 300     No new data
19 300     No new data
24 800 21 1993-95  
38 400 23 1994  
41 300 29 1987-89  
43 1000 375 1993-94  
57 2000 50 1992-94  
60 250 1100 1993-94  
65 2000     No new data
89   1700 1993  
96 650 500 1988  
97   600 1994  

 

Common Pochard

Species name
Scientific: Aythya ferina
UK: Common Pochard
DK: Taffeland

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Common Pochards occur at larger freshwater lakes, coastal lagoons, estuaries and in shallow water along coasts.

Occurrence in Denmark
Occur as a breeding, passage and winter visitor. Numbers in Denmark are highest during passage.

Trends for relevant population(s) overall and in Denmark
The northwest European winter population decreased in the period 1970-90 and is presently estimated at 350,000 birds (Scott and Rose 1996). The decrease followed a long period of westward expansion of range and population growth, which started 150 years ago (Tucker and Heath 1994). Most recently, the population shows signs of stabilisation and partial recovery (Madsen et al. 1996).

Significance of the Danish population
Up to 4% of northwest European total occur simultaneously during passage and 1-3% in winter according to Jensen (1993). The counts within SPAs (see below) suggest, though, that these figures might be underestimated, at least as regards passage.

Population trend within Danish SPAs and SPA coverage
Some of the observed decreases (e.g. in SPA no. 24) are probably a result of decreased habitat quality, but others are probably just random fluctuations.

All main staging areas are SPAs, coverage is thus high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 7000 1500 1993  
24 10000 1965 1993-95  
74 4000 740 1993-94  
78 1000 250 1996  
86 13000 1140 1995 Only part of area counted 1995
87 7-10000 12255 1995  
93 4600 3870 1987-89  

 

Tufted Duck

Species name
Scientific: Aythya fuligula
UK: Tufted Duck
DK: Troldand

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Tufted Ducks mainly rests at freshwater lakes during the day, while foraging mainly takes place nocturnally in coastal lagoons and along sheltered coasts. In harsh winters when lakes freeze over high numbers can be counted at coastal localities also during daytime.

Occurrence in Denmark
The species is a non-numerous breeding visitor and an abundant passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European winter population is thought to be increasing slowly and presently estimated at 1,000,000 birds (Scott and Rose 1996).

Significance of the Danish population
The total Danish winter population has been given as 101,000-141,000 birds (Jensen 1993) or close to 200,000 birds (Grell 1998). These figure amounts to 10-20% of northwest European total.

All major daytime roosts are covered by the SPA network. Probably most foraging areas are also situated within the SPAs, though most of these areas are not known in detail. Coverage is thus high for daytime roosts and probably also high for feeding areas.

Population trend within Danish SPAs and SPA coverage
Numbers counted in SPAs exhibit large fluctuations. These are related to winter harshness. In mild winters most birds roosts during daytime on freshwater lakes. In colder winters, when lakes freeze over the birds roost in shallow sea areas and coastal lagoons and in very cold winters some birds are forced further southwest to The Netherlands, Britain and Ireland (Scott and Rose 1996). These weather related fluctuations obscure the picture so that no clear trends are recognisable.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 9000 2400 1993  
24 9000 510 1993-95  
47 10000 1280 1991 Probably under-
estimated
64 2600 4200 1987-89  
71 23000 13140 1994  
74 7500 5450 1993-94  
78 7500 4500 1996  
81 25000 22400 1987-89  
83   22000 1987-89  
84 8000 6880 1993-94  
85   1300 1993-94 Possibly internationally important numbers occur at night
86 11000 3510 1996 Only part of area counted
87 15-20000 17650 1995  
88 12000 2600 1994  
89 19000 2475 1987-89  
93 17000 3840    
95 24000 1440 1987-89  
101 6000 720 1987-89  
105 20000 20821 1998  
109 17000 2000 1995  
111 55000 10022 1990-91  

 

(Greater) Scaup

Species name
Scientific: Aythya marila
UK: (Greater) Scaup
DK: Bjergand

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Localized, SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: II/2, III/2

General ecology and habitat requirements
Staging Greater Scaups occur mainly along coast. In daytime birds roosts in dense flocks, most foraging takes place nocturnally.

Occurrence in Denmark
The species is an occasional breeder and an abundant passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European winter population is estimated at 310,000 birds. The trend for this population is unknown as large local fluctuations and low coverage in counts before late 1980s obscure the picture. In some areas decreases are reported in others increases (Scott and Rose 1996).

Main threat is probably oil-spills. Locally mussel fisheries and hunting and associated disturbances might have an impact.

As the species is highly gregarious it is sensitive to hunting related disturbances, but the present level of hunting is probably of negligible significance for the population (Madsen et al. 1996). The species will probably benefit from the creation of new hunting and disturbance free core zones, though (Clausen et al. 1997).

Significance of the Danish population
The Danish winter population of 13,000-39,000 birds equals 4-13% of European total.

In recent years it is believed that the number of birds wintering in Denmark has decreased and the site selection of the flocks have become more unpredictable (Grell 1998). The latest available counts, from January 1997, nevertheless recorded 21,243 birds by a reduced program (Pihl et al. 1997).

Population trend within Danish SPAs and SPA coverage
Any possible overall trend inside SPAs is obscured by large year-to-year fluctuations in numbers. The coverage is intermediate, as some important areas, notably the Helnæs Bay, the south coast of Djursland and the coast near Sæby, are not SPAs.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
2 17000 9500 1993-94
12 1500 0 1991
15 3000 5000 1993-94
36   5545 1996
47 40000 2750 1988-93
64 2700 30 1987-89
71 19000 1200 1987-91
94 4000 2450 1992

 

Common Eider

Species name
Scientific: Somateria mollissima
UK: (Common) Eider
DK: Ederfugl

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2, III/2

General ecology and habitat requirements
Staging Eiders occur mainly in shallow bays, lagoon, estuaries and along sheltered coasts but also sometimes in high numbers at off-shore mussel beds.

Occurrence in Denmark
The species is common breeder (mainly resident but some are migratory) and a very abundant moulting, passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European population of ssp. mollissima has increased considerably in recent decades but might now be stabilizing at 1,735,000-2,355,000 birds. The Baltic-Danish-Dutch segment of this population is estimated at 1,350,000-1,700,000 birds (Scott and Rose 1996).

The numbers wintering in Denmark have been estimated at 538,000-797,000 birds (Jensen 1993).

Main threats are large oil spills and locally intensive mussel fisheries.

Present level of hunting probably has negligible impact on population size, as recent population trends are very similar in countries with and without an open season (Madsen et al. 1996).

Significance of the Danish population
As much as 23-46% of the total northwest European population can occur simultaneously in Danish waters. A considerably larger part of total population passes through Denmark (Clausen et al. 1997).

The birds staging in Danish waters are composed of Danish breeders, Baltic birds and birds from southern Norway and western Sweden (Jensen 1993).

Population trend within Danish SPAs and SPA coverage
Apart from random fluctuations, numbers within SPAs seems reasonably stable. In the areas affected by the construction of the Great Belt Bridge (areas no. 73, 77 and 98), numbers fell temporarily, but have since partly recovered.

The coverage is low or intermediate, as even SPAs with high densities are only small parts of larger continuous wintering areas (see Durinck et al. 1994). Hence, the present SPA network is only of limited significance for the conservation of the Danish winter population, as the species is widespread and abundant in most shallow marine waters in Denmark.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY
2     55000     77000 1993-94
9           16200 1987-91
10     80000     171000 1987-91
15     47000     70000 1993-94
31 2000 2000 62000     51000 1987-89
32     80000     98500 1987-89
36 150 150 80000     22150 1991-93
47     40000     24200 1994
52 400 400          
57     83000     60061 1995
71     45000     15183 1994
73 300 300 24000     7000 1993-94
76     20000        
77     20000     4800 1992-93
79     2000     2000  
85     700     700  
94     37000     26985 1992
96     10000     12400 1987-89
98     36000     9400 1992-93
102     34000     30000 1996
110 5500 5500   7000 7000   1993-96

 

Long Tailed Duck

Species name
Scientific: Clangula hyemalis
UK: Long-tailed Duck
DK: Havlit

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Long-tailed Ducks are less restricted to shallow water than most other seaducks. It mainly occurs in offshore areas with a depth between 10 and 35 m metres (Durinck et al. 1994)..

Occurrence in Denmark
The species is a winter visitor to Denmark.

Trends for relevant population(s) overall and in Denmark
Trends for the northwest European winter population are not known as it has recently been realised that this population is considerably larger than previously believed. Following extensive surveys in the Baltic, Durinck et al. (1994) concluded that the population probably numbered 4,700,000 birds, while Rose and Scott (1996) estimate 4,600,000 birds. Numbers in the breeding areas in Greenland, Iceland, Scandinavia and Siberia are thought to be stable (Scott and Rose 1996), though, so probably the winter population is also reasonably stable.

Meranian Bay and numbers in Danish waters rise sharply.

Oil-spills are probably main threat.

As the species mainly occurs dispersed and offshore it is not very susceptible to hunting related disturbances. The present level of hunting is of negligible significance for the population (Madsen et al. 1996).

Significance of the Danish population
Danish waters holds from 165,000 birds in mild winters up to 425,000 birds in cold winters (Jensen 1993). In cold winters, thus, up 9% of total population is in Danish waters.

Population trend within Danish SPAs and SPA coverage
No Danish SPA holds internationally important numbers on a regular basis. The most important areas in Danish waters are further offshore than most SPAs and are situated south of Lolland, south and east of Falster and at Rønne Banke south of Bornholm. The coverage for this species is thus low and the present SPA network is of negligible significance for the Danish winter population. Danish waters (including non-SPA areas) are only of secondary importance for this species compared to other parts of the Baltic.

The species is included here as the aggregation 6,000 birds counted in DSPA no. 71 was at designation time believed to be of international importance. As total population is now known to be very large (see above), aggregations must meet the 20,000 waterbirds criterion to qualify.

 

Common Scoter / Black Scoter

Species name
Scientific: Melanitta nigra
UK: Common Scoter / Black Scoter
DK: Sortand

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2, III/2

General ecology and habitat requirements
Moulting and wintering Common Scoters occur mainly in off-shore areas with shallow water (mostly 5-15 metres).

Occurrence in Denmark
The species is a visitor occurring in high numbers in Danish waters both during moult and in winter.

Trends for relevant population(s) overall and in Denmark
The total population breeding from western Siberia to Scotland and Iceland and wintering from northwest Europe to Mauritania has recently been estimated at 1,600,000 birds (Scott and Rose 1996).

As the species mainly occur offshore and has a large population the present level of hunting and associated disturbances are probably of negligible significance for the population (Madsen et al. 1996).

Significance of the Danish population
Up to more than 940,000 birds can occur simultaneously in Danish waters (Durinck et al. 1994); this equals 59% of total population.

Population trend within Danish SPAs and SPA coverage
Numbers inside SPAs are highly fluctuating. The main reason for this is that most SPAs are only parts of larger continuous wintering areas. The internationally most important area is a 4,690 km2 area in northern Kattegat which on average holds 494,100 birds. Especially in cold winters also the eastern German Bight (= off-shore parts of the Wadden Sea) can hold very high numbers (190,000 birds) (Durinck et al. 1994). Of the area in northern Kattegat, only small parts are included in the SPA network (areas no. 2,9,10,15 and 32). The situation in the German Bight is very similar only a small part of the area is included in SPA no. 60. The coverage for this species is accordingly low.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
2 38000 18000 1993-94
10 80000 50000 1987-91
15 10000 28000 1993-94
31 9000 9940 1993-94
32 20000 8800 1987-89
36 8000    
57 200000 176700 1987-89
73 5000    
76 4200    
77 6000    
94 15000 16300 1988-93
96 5000    
102 4000    

 

Velvet Scoter

Species name
Scientific: Melanitta fusca
UK: Velvet Scoter
DK: Fløjlsand

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Localized, SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Winters in often large and dense flocks mostly offshore in areas with a depth of 10-30 metres.

Occurrence in Denmark
The species is a moulting and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European winter population is believed to be reasonably stable (Scott and Rose 1996) though trends are not well known as the high numbers occurring in several areas in the Baltic were only recently discovered. As a result of these discoveries the estimated total of this population has been changed from 250,000 to 1,000,000 (Durinck et al 1994, Scott and Rose 1996).

Present size of the Danish population is only known within broad limits. The Danish winter population was recently estimated at 150,000 birds in the mid-1990Žs with an unknown trend in the period 1978-94 (Asbirk et al. 1997). Grell (1998) gives the wintering population as 100,000 birds while Jensen (1993) estimates 125,000.

Main threat toward the population is oil spills (Tucker and Heath 1994).

The present level of hunting is probably of negligible significance for the population (Madsen et al. 1996).

Significance of the Danish population
Most of the birds wintering in the Baltic are outside Danish waters, but 10-15% can be within; most of which occur in northern Kattegat.

Population trend within Danish SPAs and SPA coverage
The numbers recorded within Danish SPAs are very fluctuating and generally not of international importance (1% = 10,000). The most important area within Danish waters is in northern Kattegat. Only minor parts of this large area are covered by SPAs no. 2,9,10,15 and 32. Consequently, the coverage for the species is low.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
2 20000    
10 11000   1983
12 16000 0 1991
15 20000 11000 1993-94
36 3000 8 1995
47 5000 2 1991
71 4000    
73 600    
76 2000 1 1988-93
77 600    
94 12000 3250 1988-93
96 3000 723 1987-89

 

Common Goldeneye

Species name
Scientific: Bucephala clangula
UK: (Common) Goldeneye
DK: Hvinand

Red-list and SPEC status DK/Europe/World
DK: Rare (breeding) and Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Goldeneyes occur both in fresh, brackish and salt water. In cold winters when lakes freeze over nearly all are at sea (Jensen 1993).

Goldeneyes rarely form large flocks but are usually scattered along the coast in small flocks. In mild winters, it is restricted to areas of less than 10 metres water depth, close to shore. In hard winters often move offshore where they concentrate in small ice-free waters (Durinck et al. 1994).

Occurrence in Denmark
The species is a newly established breeder and a common visitor during moult and in winter.

Trends for relevant population(s) overall and in Denmark
The winter population in northwest and central Europe is increasing in northwest and stable in central Europe. The total for this population is estimated at 300,000 birds (Scott and Rose 1996).

The present level of hunting probably has a negligible impact on the population and because the species is not gregarious, it is not very sensitive to hunting related disturbances (Madsen et al. 1996).

Significance of the Danish population
Up to 65,000 Goldeneyes occur simultaneously in Denmark (Clausen et al. 1997). This equals 22% of total northwest and central European winter population.

Population trend within Danish SPAs and SPA coverage
Most observed changes within SPAs are probably random fluctuations. In some cases (notably in area no. 12), though, observed decreases are probably a result of decreased habitat quality.

The coverage is high, most important wintering areas (see Durinck et al. 1994) for the species are included in the DSPA network. A few important areas along the coast of east and especially southeast Jutland are not SPAs, though.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 3500 590 1993  
2 4000     No new data
3 3000 150 1994  
12 12000 759 1994-96  
14   3370 1993-94  
15   3110 1993-94  
24 7000 2938 1993-93  
26 1500 9500 1993  
27 4000 330 1994  
28 3000 0 1991  
36 1 3444 1996  
40 15000 1200 1995  
43 3200 750 1993-94  
47 5000 3000 1988-93  
64 400 370 1987-89  
71 4000 3050 1994  
83 200 810 1987-89  
85 5000 1350 1993-94  
86 2000 75 1995 Only part of area counted 1995
89 5000 3430 1991  
97 2000 2200 1994  
102 2000 500 1996  
105 2500 5127 1998  

 

Smew

Species name
Scientific: Mergus albellus (or Mergellus albellus)
UK: Smew
DK: Lille skallesluger

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Wintering Smews mainly occur in coastal lagoons and other in-shore areas, though small numbers winter in freshwater lakes in mild winters.

Occurrence in Denmark
The species is winter visitor to Denmark.

Trends for relevant population(s) overall and in Denmark
Present trend in the northwest and central European population is unclear. Estimate of total numbers has recently been revised from 15,000 to 25,000-30,000 birds (Scott and Rose 1996). The new estimate is largely the result of the discovery of very large concentrations in coastal lagoons on the Baltic coast of Poland and Germany (Durinck et al. 1994).

Main threat on a European basis is currently believed to be predation in breeding areas by the introduced American Mink (Mustela vison). Large population decreases and range contractions from mid 19th century to around 1970 are believed to be the result of deforestation in river valleys, forestry intensification and dam building (Tucker and Heath1994).

Significance of the Danish population
Up to 1,000 birds occur simultaneously in Danish waters, in mild winters the numbers are lower (Jensen 1993). This only equals 3–4% of total northwest and central European total.

Population trend within Danish SPAs and SPA coverage
Numbers inside SPAs fluctuate markedly in relation to winter harshness. The coverage is high, as all areas shown as nationally important for the species in Jensen (1993) are at least partly SPAs (no. 24, 81, 84, 96 and 111). The only area, which regularly supports internationally important numbers, is SPA no. 111. Very high numbers also occur, although more irregularly (up to 800 reported), in SPA 84 and adjacent areas. The coverage is high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
111 400 184 1993

 

Red-Breasted Merganser

Species name
Scientific: Mergus serrator
UK: Red-breasted Merganser
DK: Toppet Skallesluger

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Winter gregariously in flocks up to a few hundred birds in sea areas with a depth of less than 15 metres (Durinck et al. 1994). In autumn highly concentrated in a few areas, but more dispersed in winter (Grell 1998).

Occurrence in Denmark
The species is a common breeder and an abundant passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest and central European winter population is believed to be largely stable and is currently estimated at 125,000 birds (Scott and Rose 1996).

Significance of the Danish population
Up to 27,660 birds or 22% of total northwest and central European population have been registered in Danish waters during passage (late autumn). In winter numbers counted are in the range 8,300-15,000 birds (Jensen 1993). The numbers counted during moult (August) fluctuated in the late 1980s (Jensen 1993) and these aggregations seem to have disappeared almost totally in the 1990s (Grell 1998).

Present level of hunting and associated disturbances are probably of negligible significance for the population (Madsen et al. 1996).

Population trend within Danish SPAs and SPA coverage
Numbers counted in Danish SPAs are in many instances lower in recent years than at time of designation. Most of the observed changes are probably random fluctuations, but some may be the result of decreased habitat quality (e.g. at SPAs no. 12 and 14).

The coverage is high as most areas of high densities according to Durinck et al. (1994) are within the SPA network.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 1000 1550 1993  
2 1000 105 1993-94  
10 2000   1983  
12 10000 1800 1994-96  
13 2000 1560 1994  
14 10000 73 1993-94  
23 700 50 1996  
27 2000 304 1994  
28 2000 330 1988  
38 500 15 1996  
40 1000 400 1995  
47 4500 351 1995 Only part of area counted 1995
57 2000 50 1992-94  
64 100 4 1991  
71 1000 4520 1994  
75 1000 400 1993-94  
81   1704 1987-89  
83   1060 1987-89 Data incomplete
84 2000 270 1993-94  
85 4000 3525 1987-89  
88 500 595 1996  
89 1200 1650 1987-89  
102 1000     No new data
110 950 298 1994  

 

Goosander

Species name
Scientific: Mergus merganser
UK: Goosander
DK: Stor skallesluger

Red-list and SPEC status DK/Europe/World
DK: Rare (breeding)
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Goosanders occur both in fresh and salt water. Main areas are larger lakes and coastal lagoons. Highest numbers at coastal localities occur in cold winters when freezing displaces birds wintering in lakes.

Occurrence in Denmark
Rare resident and common passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The northwest European population is thought to be stable or slightly increasing. The combined northwest and central European population is estimated to number 200,000 birds (Scott and Rose 1996).

No major threats towards the population are known. Locally (SPA no. 12) decreased habitat quality as a result of eutrophication might have had an impact (Jensen 1995).

Present level of hunting and associated disturbances believed to be of negligible significance for the population (Madsen et al. 1996).

Significance of the Danish population
Up to 21,400 birds have been recorded simultaneously in Danish coastal areas (Jensen 1993). This equals 10-11% of total northwest and central European winter population.

Population trend within Danish SPAs and SPA coverage
Numbers counted in the Danish SPAs fluctuates, mostly dependent on winter harshness. At a few sites (e.g. no 12) numbers have probably decreased as a result of decreased habitat quality.

The coverage is high, almost all coastal sites regularly supporting high densities according to Durinck et al. (1994) are within the SPA network and most important freshwater sites are likewise covered.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 1500 410 1993  
12 2000 161 1994-96  
14 3000 2780 1993-94  
15 1000 230 1995-96  
24 600      
28 4000 200 1988  
29 2000 2046 1994  
30 1000 34 1993-94  
31 1000 146 1993  
33 800 440 1995  
35 1200 100-200 1993-94  
36 1000 75 1996  
38 2500 29 1996  
40 1000 310 1995  
43 5000 625 1993-94  
57 3000 150 1992-94  
60 1700 92 1991  
75 1000 900 1993-94  
81   2610 1987-89  
84 1500 62 1993-94  
85 1500 233 1993-94  
86 2000 132 1994 Only part of area counted
87 1200 1140 1995  
89 2000 450 1991  
102 1500     No new data
105 2500 5801 1996  
106 2600 10000-12000 1998  
109 800 150 1995  
111 1000 505 1995  

 

Honey Buzzard

Species name
Scientific: Pernis apivorus
UK: Honey Buzzard
DK: Hvepsevåge

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
The Honey Buzzard breeds in old broad-leaved forests near more open areas - often wetlands or natural grasslands - suitable for foraging.

Occurrence in Denmark
The species is a breeding visitor and a passage visitor from other parts of Scandinavia, probably mainly Sweden.

Trends in Danish population - long-term and 1983-1998
The long-term trend in the Danish population is not known with certainty, but it is believed that no major changes have taken place recently (Jørgensen 1989). Earlier the population estimates were considerably lower (200-300 pairs, Dybbro 1976) than what is now believed to be the case (650 pairs, Jørgensen 1989 and Grell 1998). In all likelihood, this reflects increased knowledge rather than any real growth in the population.

Main threats toward the population are intensified forestry (including conversion of broad-leaved forest to conifer plantations, shorter cycles and drainage of wetlands within forest areas) and hunting during migration mainly in Malta and Italy (Grell 1998).

Current efforts restoring smaller areas of pastoral woodlands are probably beneficial for the species.

Significance of the Danish population
The Danish population is only a small part (well below 1%) of the total European population that was recently estimated 111,200-148,672 pairs plus (Hagemeijer and Blair 1997).

Population trend within SPAs, SPA coverage and significance of SPAs for species

The estimated population within the DSPAs seems a bit higher than at designation time, but as is the case with the total Danish population this probably only reflects better knowledge. Conceivably the population in the SPAs is still somewhat under-estimated as there is a lack of recent data from some former breeding areas, notably DSPA no. 45.

The coverage is low, less than 10% of the Danish population breed within the SPAs. This is a result of the species’ dispersed breeding distribution.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP Min. BP Now Max. BP NDY Comments
04 1 1 4 5 1993  
33     2 2 1993-94  
34 1 1 2 4 1993-94  
37     1 1 1993-94  
45 5 6       No new data
58     0 1 1989  
59 4 4     1983 No new data, but probably still breeds
61       0 1 1996
66 2 3 0 1 1997  
68 5 5     1983 No new data, but probably still breeds
70     0 1 1993-94  
74 1 1 4 4 1994  
80 1 1 2 3 1995  
81     2 2 1994-95  
87     6 6 1986-90  
90 1 1 1 1 1994  
91     1 2 1993-94  
92 P P       No new data
107 4 6 3 5 1995  
108 8 10 9 8 1993-94  
Total 34 40 37 47    

 

Red Kite

Species name
Scientific: Milvus milvus
UK: Red Kite
DK: Rød glente

Red-list and SPEC status DK/Europe/World
DK: Rare
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
The species prefer open country with scattered woodlands. As carrion forms a significant part of the diet, several pairs breed near highways providing roadkills.

Occurrence in Denmark
Occurs as a breeding visitor and as a passage visitor from the population in South Sweden.

Trends in Danish population - long-term and 1983-1998
The species was a widespread and common breeder until the start of intense persecution of almost all raptors, which followed the introduction of pheasantry around 1880. Total eradication followed during the first decade of the century. The species became totally protected in 1922, but despite scattered breeding attempts during the period 1910-1970, actual re-colonization did not occur until the 1970s. Since then, the population has been slowly increasing. Despite rapid increase in southern Sweden, the Danish population is still concentrated in southeast Jutland. Comparison with neighbouring countries yield estimates of potential for at least 400-500 pairs within Denmark (Grell 1989). This potential is far from fulfilled and illegal persecution by shooting or poisoned bait probably plays a larger role for the population trend of the present species than for any other Danish raptor, possibly excepting Goshawk (Accipiter gentilis). Though few cases are proven, the problem is supposed to be more widespread (Grell 1998).

Significance of Danish population
Presently the Danish population is only a very small part of total European population, which was recently estimated at 17,395-28,235 pairs Hagemeijer and Blair 1997). As mentioned above there are considerable possibilities for a more significant Danish population in the future.

Population trend within Danish SPAs and SPA coverage
As can be seen only a very small part of the Danish population breeds within SPAs. The distribution map in Grell 1998 indicates that the table above is not complete as a few pairs seem to inhabit other SPAs, than those included here, no details are known. Probably the coverage is in the 10-20% range. Most Danish pairs has been established after the designation of the DSPAs.

Some SPAs are also used as short-term staging areas during migration periods.

The low coverage reflects the dispersed nature of the breeding population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY
80     2     3 1997
82           14 1993
83           7 1993-95
90           3 1997
Total     2     3  

 

White-tailed Eagle

Species name
Scientific: Haliaetus albicilla
UK: White-tailed Eagle
DK: Havørn

Red-list and SPEC status DK/Europe/World
DK: Rare, i*
Europe: Rare, SPEC 3
World: Lower Risk (near threatened)
Annexes: I

General ecology and habitat requirements
In Denmark, the species breeds in older broad-leaved forest near large lakes or fjords. In winter mainly in coastal areas.

Occurrence in Denmark
The species is a breeding resident and a passage and winter visitor from the rest of Scandinavia.

Trends in Danish population - long-term and 1983-1998
The European population has been reduced over centuries through shooting, poisoning and nest robbing. Loss and degradation of wetlands, disturbance at nest sites and intensification of forestry are other threats (Tucker and Heath 1994).The original Danish population was persecuted to extinction, the last breeding occurred in 1911. Around 50 breeding sites are known from the period 1850-1899 (Grell 1998).

There have been sporadic breeding attempts during the years 1952-60 (3 sites) and 1979-1980 of which a few where successful, but actual re-colonisation did not occur until 1996 where two pairs bred successfully, each fledging one young. In 1997, three pairs made breeding attempts (two successful) and in 1998, those three pairs and one additional pair were all successful fledging a total of 6 young (Grell 1998, J. J. Madsen pers. com.). In 1999, 6 pairs made breeding attempts, 5 were successful, and 7 young fledged (M. Grell pers. com.). An additional 2-3 pairs are under establishment.

The re-colonisation is believed mainly to be the result of the population increase in Northern Germany as at least some of the Danish breeders are colour-ringed as nestlings there (Grell 1998).

Until now, disturbance at nest sites has been avoided by the creation of no admittance zones near nests. With the increasing number of breeding pairs this might in the near future not be possible at all sites.

Approximately 20-40 birds probably mostly from Sweden winter in Denmark, mainly at coastal localities in the eastern part of the country. The number is highest in harsh winters (Grell 1998).

Significance of the Danish population
The Danish population is still very small compared to the European total of 3,292-3,550 pairs (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
All Danish pairs breed in connection to SPAs, though some of the actual nest sites are just outside the areas. The mentioned 2-3 pairs under establishment are all within SPAs too, so the coverage will probably remain near 100% in the near future. Appropriate management of SPAs is thus of the highest importance for the continued success of the species in Denmark.

Most of the wintering visitors are also within SPAs.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Com-
ments
7           3 1997  
13           P 1997  
15           1 1997  
32           2 1997  
33           2 1995  
35           2 1993-94  
36           1 1996  
37           3 1997?  
43           1-2 1998  
47       1 1   1998 First breeding 1996
55           2 1996  
58       1 1   1996  
60           1 1997  
74           4 1998  
76           1 1995  
78       1 1   1998  
80           2 1997  
81       1 1 5 1999  
82           12 1997  
83           1 1997  
85           1 1997  
87       1 1   1998 First breeding success 1997
89       1 1 3 1999  
93           6 1997  
96           >2 1998  
97           1    
99           2 1997  
105           4-7 1997  
106           2 1998  
Total       6 6      

 

Marsh Harrier

Species name
Scientific: Circus aeruginosus
UK: Marsh Harrier
DK: Rørhøg

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Marsh Harriers breed in reed beds and forages both in wetlands, other open natural habitats and in cultivated areas (Grell 1998).

Occurrence in Denmark
The species is a breeding visitor and a passage visitor from other parts of Scandinavia, probably mainly Sweden (Grell 1998).

Trends in Danish population - long-term and 1983-1998
In the mid 19th century the species was reasonably common in most parts of the country. Intensive persecution decimated the population and at the turn of the century, it was almost eradicated. After 1922, it was protected during the breeding season and the species partly recovered only to be re-decimated by habitat destruction and environmental pollutants in the 1950s and 60s. As a result of the total protection of Danish raptors in 1967, the somewhat improved protection on the migration route and the ban on some of the most harmful pesticides the Danish population has been steadily increasing from 50-80 pairs in the late 1960s to the present population of approximately 650 pairs (Grell 1998).

Presently the main threat toward the population is probably hunting during migration, mainly in Italy and Malta (Grell 1998). In Denmark disturbance at nest sites and inappropriate water level management can play a role at least locally.

Significance of the Danish population
The European population is estimated at 50,955-74,675 pairs and the Danish population is thus approximately 1% of the total.

Population trend within Danish SPAs and SPA coverage
The populations within the SPAs seem to have followed the general trend in the Danish population quite closely. The population within SPAs is probably a bit underestimated as data for some areas are lacking. The coverage is probably in the 30-40% range. The remaining part of the Danish population mainly occurs dispersed at smaller localities harbouring single or a few pairs.

The DSPA network protects key populations of high density, so despite the majority of the population nesting outside the areas, appropriate management of the DSPAs is of high importance for the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Com-
ments
01       2 2   1993  
07       2 2   1994  
13 10 10   10 10   1994  
15       4 4   1995-96  
20 1 1   1 1   1995  
22       1 1   1993-94 Breeding data are from 1987-88
23 1 1   0 2 7 1996 Breeding data are from 1987-88
24           23 1993-95  
27       1 1   1995  
30       1 1   1988-89  
35 1 1   0 1   1994  
36       1 1   1996  
37 1 1   2 2   1995  
38 3 4   4 4   1993-96  
39       0 0   1998  
41 1 1   3 4   1995  
43 11 11   9 9   1993-94  
44 1 1   0 0   1993  
47 1 1   3 4   1994-96  
49       3 3   1991-94  
51     20 2 4   1993-94  
52 1 1   2 2   1991  
53 3 3   3 4   1993-94  
55       P P 9 1996  
56 1 2   1 2   1993-94  
57 3 4   4 4   1991  
58 2 3   1 1   1989  
60 33 35   20 20   1996  
61 1 2   0 0   1993-94  
62 1 2   0 0   1994  
63 1 2   0 0   1989  
65 1 1   1 3   1994  
67       2 2   1994  
68 1 4   0 0   1993-94  
69 1 1   0 0   1993-94  
70 1 3   0 1   1993-94  
71 7 7   10 15   1996  
72 1 1   3 5   1995  
75 2 2   P P   1994  
76 3 3   2 2   1994  
78 2 2   2 2   1996  
80       8 9   1997  
81       5 5   1995  
82 3 5   1 1   1993  
83 4 5   5 5   1993-96  
84       3 3   1985-89  
85 1 1   3 3   1997-98  
86 1 1   1 1   1985-89 No new data
87 15 30   35 35   1995  
88       1 1   1996  
89 4 4   4 4   1993-96  
91 1 3   8 10   1993-95  
93     1 0 0   1993-94  
95 2 6   6 8   1989  
96 5 10 1 9 9   1987-89  
97       1 1   1994  
99 1 2           No new data
100 2 3   4 5     1993-94
101 4 4       4 1997 No new breeding data
103 1 1   0 1   1996  
104 1 1   0 1   1995  
106 2 3           No new data
109 1 1   0 1   1995 Breeds most years
110       5 6 10 1997  
111 1 3   1 1   1995  
Total 145 194   201 230      

 

Hen harrier

Species name
Scientific: Circus cyaneus
UK: Hen Harrier
DK: Blå kærhøg

Red-list and SPEC status DK/Europe/World
DK: Rare
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
The species is known to breed in a wide variety of open and semi-open habitats (Tucker and Heath 1994). The few Danish pairs have been found in meadows, coastal marshes, moors and heathlands (Grell 1998). Foraging mainly takes place in areas of tall grass (Jørgensen 1989).

Occurrence in Denmark
It is not known whether the very few Danish breeders are visitors or resident. The species also occur as passage and winter visitor from the substantial populations in the rest of Scandinavia.

Trends in Danish population - long-term and 1983-1998
The species has always been a scarce and irregular breeder in Denmark with breeding records only from the years around 1900, the 1930s, 1943, 1971-82 and since 1992 (Grell 1998). The re-colonization in 1992 represents an extension of the Dutch Waddensea population, which has been increasing since the 1950s (Hagemeijer and Blair 1997).

Main threat at a European level is widespread loss of breeding habitat (Tucker and Heath 1994). The loss mainly occurs through agricultural intensification, but can also occur through abandonment or afforestation.

Significance of the Danish population
The Danish breeding population is very small (2-5 pairs, Grell 1998) compared to the European total of 23,332-30,840 pairs (Hagemeijer and Blair 1997). Internationally Denmark is more important for staging and wintering birds from other parts of Scandinavia. In 1989 the Danish winter population was estimated at 700-900 birds but believed to be declining (Jørgensen 1989).

Population trend within Danish SPAs and SPA coverage
For the small breeding population, the coverage is probably 100% at least in most years.

The SPAs also hold a very substantial part of the birds staging and wintering in Denmark. The knowledge of the total number of staging birds in Denmark and within the individual SPAs is not accurate enough, though, to allow a more detailed analysis.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Com-
ments
1           5 1998  
7           3 1993-94  
13           20 1997  
14           1 1996  
20           10 1996  
23           4 1994  
33           3 1993-95  
35           3 1993-94  
37       1 1 4 1993-94  
38           8 1997  
41           7 1997  
43           16 1998  
44           2 1996  
49       3 3   1996  
51     10          
55           4 1996  
65       1 1   1997 Breeds some years
67     10          
71           2 1997  
72           4 1997  
76           2 1997  
80           1 1997  
81           >5 1983-98  
85           1 1997  
89           >10    
91           8 1994  
93           2 1997  
96           >5 1998  
97           1    
110           6 1996  
111           4 1997 Formerly much higher numbers staging

 

Montague’s Harrier

Species name
Scientific: Circus pygargus
UK: MontagueŽs Harrier
DK: Hedehøg

Red-list and SPEC status DK/Europe/World
DK: Vulnerable
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
In Denmark the species mainly breed in heathland moors (earlier the main habitat now only a few pairs), reedbeds and increasingly in cereal fields, roughly half the Danish pairs now nest in such habitats (Grell 1998).

Occurrence in Denmark
Occurs as a breeding visitor and as passage visitor from the Swedish (and possible Finnish) population (Grell 1998).

Trends in Danish population - long-term and 1983-1998
The species first colonized Denmark at the turn of the century and the colonization was followed by a period of rapid range expansion and population growth. The population reached a maximum of 350-400 pairs around 1940. Hereafter a rapid decrease followed and a thorough survey of former breeding localities in the years 1968-1972 showed that the population was reduced to a mere 30-40 pairs (Jørgensen 1989) primarily as a result of drainage and agricultural intensification. The population size has fluctuated near that level since with 35-50 pairs in 1993-96 (Grell 1998).

Pairs nesting in cereals often loose clutch during harvest. An intensive monitoring program of the Danish population by BirdLife Denmark and the Danish Animal Protection Society and dialogue with involved farmers seems to have minimised this problem. No clutches have been lost as result of harvest since start of the project in 1994 (Grell 1998).

Significance of the Danish population
The Danish population is only of marginal significance compared to the estimated total European population of 26,976-39,610 pairs (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
Although the overall Danish population has been roughly stable apart from year-to-year fluctuations, the population within SPAs seems to have decreased since designation. This trend is probably associated with the breeding habitat shift of part of the Danish population to cereal fields. Possible reasons for the decline in some areas could be increased disturbance from recreational activities and inappropriate management of water levels at some sites.

The decrease within SPAs also results in a decreased coverage. As the reasons for this shift are unclear, it is uncertain whether it reflects inappropriate management of some sites or it is caused by other factors. Presently the coverage is probably in the 50-70% range. Despite the decrease in coverage the most important Danish populations still breed within SPAs and some pairs now breeding outside SPAs still utilise SPAs as foraging areas.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP Min. BP Now Max. BP NDY Comments
1     1 1 1998  
13     1 1 1998 No breeding success
43     0 0 1993-94  
49     3 3 1996  
51     1 2 1993-94  
53            
56 1 1 0 0 1993-94  
57 7 10 2 2 1991  
60 7 18 3 3 1996  
61 1 2 0 0 1993-94  
62 1 1 1 1 1994  
63 3 5 0 0 1989  
65 P P 9 9 1994  
67     4 4 1994  
69 3 4 0 0 1993-94  
70 2 3 0 0 1993-94  
Total 26 45 25 26    

 

Species accounts, part 2

 

Black Grouse

Species name
Scientific: Tetrao tetrix
UK: Black Grouse
DK: Urfugl

Red-list and SPEC status DK/Europe/World
DK: Endangered
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: I (continental ssp. tetrix only), II/2

General ecology and habitat requirements
In Denmark, the species occurred in a mosaic landscape of lowland heaths often near bogs and areas of low-intensity agriculture.

Occurrence in Denmark
The species was a breeding resident.

Trends in Danish population - long-term and 1983-1998
In the 19th century, the species was widespread and abundant in Jutland. The species is not indigenous to the Danish isles, though some short-lived introduced populations were present last century.

Habitat destruction and fragmentation through afforestation and agricultural intensification are main threats at a European level. In some areas, eutrophication through airborne nitrogen deposition is also considered a major threat (Tucker and Heath 1994).

In Denmark, the general conversion of heathland to agriculture or forestry and the agricultural intensification during the last 150 years resulted in major declines and the final extinction of the population.

In 1942 the population was estimated at 2400 birds, 1100 in 1966, 420 in 1973, 35 in 1991 and finally in 1997 a mere 3 (three) birds were left (Dybbro 1976, Grell 1998).

The various conservation and recovery measures proposed and implemented failed to save the population, due to lack of in depth knowledge of the problems and because of conflicting interests.

Significance of the Danish population
None.

The total European population is estimated at 678,136-1,878,889 "pairs". The species is declining in most of Europe with the exception of countries such as Sweden, where specialised management is applied. The Danish population belonged to ssp. tetrix of which only small, scattered populations remain (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
Within the DSPA network, the species only occurred in the two areas above at designation time. The species had disappeared from other formerly important areas such as SPA no. 50 less than a decade before designation.

The last two areas, which were of importance for the species (Vind Hede and Kongenshus Hede), are not SPAs, but Vind Hede is a proposed SCI (designated for the habitat 4030 European dry heaths).

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
37 P P     0 1 1998 1998 occurrence probably the result of illegal release of bird(s) of unknown origin.
46 P P     0 0   Last seen 1992

 

Spotted Crake

Species name
Scientific: Porzana porzana
UK: Spotted Crake
DK: Plettet rørvagtel

Red-list and SPEC status DK/Europe/World
DK: Vulnerable
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Occur in swamps and fens and overgrown edges of lakes and rivers (Tucker and Heath 1994). Habitat must include areas of very shallow, not oligotrophic water interspersed with ample stands of low plant cover, rich in invertebrate food (Snow and Perrins 1998). The species has a particular preference for areas dominated by sedges (Grell 1998).

Occurrence in Denmark
The species is a breeding visitor and a passage visitor probably mainly from Sweden.

Trends in Danish population - long-term and 1983-1998
The species was considerably more abundant and widespread in Denmark during the 19th century than today. It decreased markedly during the first decades of the 20th century. The population was estimated at 43-115 "pairs" (i.e. singing males) during the years 1960-69. There are considerable year-to-year variations in numbers possibly related to prevailing weather in the spring migration period. During the years 1993-96 the number of singing males recorded per year varied from 22 to 50 possibly 61 (Grell 1998).

Main threats toward the population are habitat destruction and degradation especially through inappropriate water level management, eutrophication and too high vegetation as result of abandonment (Stoltze and Pihl 1998).

Significance of the Danish population
The Danish population is only a very small part of the European total, which was estimated at 58,786-167,083 pairs by (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
Although numbers recorded in the DSPAs are higher in recent years than at designation, this almost certainly reflects better coverage rather than any real population increase.

The coverage is probably above 80% and very few regular breeding sites are not SPAs. The conditions within the SPAs thus are of high importance for the future of the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
13       13 13   1994  
20       2 2   1995  
29       0 1   1987-89  
41       0 1   1988-89  
43       0 0   1993-94  
58 0 1   0 0   1989  
60 8 8   2 2   1996 Higher numbers in most years
65       0 1   1988  
69 1 1   0 0   1993-94  
72       1 1   1996  
80       5 5   1996  
82       1 1   1993  
111       0 1   1997  
Total 9 10   24 28      

 

Corncrake

Species name
Scientific: Crex crex
UK: Corncrake
DK: Engsnarre

Red-list and SPEC status DK/Europe/World
DK: Extinct*
Europe: Vulnerable, SPEC 1
World: Vulnerable
Annexes: I

General ecology and habitat requirements
Breeds in extensively used hay meadows, drier parts of fens, clover and rape fields and sometimes bushy forest clearings (Tucker and Heath 1994). In Denmark, the species persisted most recently in meadows in river valleys.

Occurrence in Denmark
The species is a breeding visitor and a passage visitor probably mainly from Sweden.

Trends in Danish population - long-term and 1983-1998
Main threats are habitat loss through drainage and agricultural intensification. Both young and adults are also directly killed during harvest (Tucker and Heath 1994).

Thorup (1999) summarises data from several surveys and lists five demands, which have to be met for an area to support breeding Corncrakes:

  1. Habitat: Moist hay meadows, not fertilised, fringed by areas with higher vegetation (e.g. reeds, nettles, reed grass, iris)
  2. Soil moistness: Moist or wet soil is of impor tance, at least in order to avoid intensification of use and to secure late mowing.
  3. Mowing: Mowing after 1 August; trapping birds in the middle of the field should be avoided.
  4. Grazing: Breeds mainly in ungrazed meadows. Aftergrazing and low cattle density is tolerated.
  5. Other: A social breeder. Corncrake units should consist of 200-400 ha as minimum.

The species was formerly a both abundant and widespread Danish bird. Løppenthin (1967) states that during the 1860s and 1870s it was probably more abundant than Coot (Fulica atra) and Moorhen (Gallinula chloropus). By detailed analysis of land use data and comparison with density data from recent strongholds (Belarus and the Baltic) Thorup (1999) estimates the Danish population 1850-1920 to 10,000-20,000 pairs.

The dramatic decrease of the Danish population probably started during the last part of the 19th century (Grell 1998), though killing during harvest attributed to changes in cropping practice was noted as early as 1852 (Kjærbølling 1852).

Since 1960 only three sites, all river valleys, seems to have been regular breeding localities: the Skjern Å valley, the Varde Å valley and the Ryå valley (Grell 1998). In the last two areas the disappearance of the species is probably related to highly intensified production of grass pellets with large inputs of fertiliser and several croppings a year. These problems are sought remedied by restoration plans (see below).

All the three last strongholds (the Ryå, Skjern and Varde River valleys) are targeted for habitat restoration efforts. The current efforts in the Ryå valley and even the ambitiously named "Project Corncrake" in the Varde Å valley are, though, possibly insufficient to create real "recovery areas" as Thorup (1999) states that four of the "five demands" (see above) are unmet. Some of the remaining problems, though, are expected be solved during the practical implementation of the project (Jørn Jensen, NFNA pers. com.).

In the Skjern Å valley, the project’s eventual impact on Corncrakes is yet unclear.

In the spring 1999, there was a large influx to Denmark and more than a hundred calling males were recorded. Most were recorded in areas where chances of breeding success are very slim.

Significance of Danish population
Presently Denmark is of very limited if any significance for the species. There remains some chance, though, that various habitat restoration efforts notably the large project in the Skjern Å river valley might allow the re-establishment of the species as a regular Danish breeder, or if still that, a substantial increase in the population.

The total European population is estimated at 97,470-196,920 (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
The population trend within and outside SPAs have been much the same. The coverage of the small, hypothetical Danish population is probably 0%.

The only SPA with some potential for becoming of importance for the species is probably the former breeding stronghold Varde Å valley in area no. 49. This river is the only larger stream in Western Jutland, which has not been canalised, but the area is presently not managed in a manner compatible with the requirements of breeding Corncrakes.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
44 P P     0   1993  
47 P P     0   1988  
70       0 1   1994 Possibly the pair bred on German side of border
Total 2 ?   0 1      

 

(Common) Coot

Species name
Scientific: Fulica atra
UK: (Common) Coot
DK: Blishøne

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/1, III/2

General ecology and habitat requirements
Staging Coots occur highly gregariously in very shallow fresh or brackish waters (generally, less than 3 metres water depth) along sheltered coast, in lagoons, estuaries and lakes with abundant supply of macrophytes.

Occurrence in Denmark
Birds breeding north and northeast of Denmark are migratory and winter in suitable coastal areas. The Danish breeders are mainly resident or dispersive, though some are displaced during cold winters.

Trends for relevant population(s) overall and in Denmark
The northwest European winter population is estimated at 1,500,000 birds (Rose and Scott 1997). Up to 163,900 winter in Danish waters with the highest numbers in mild winters and the lowest numbers at the end of a string of cold winters (Jensen 1993). The Danish winter population was slightly decreasing 1978-94 (Asbirk et al. 1997). While it is certain that staging numbers decreased between 1968-73 and 1987-89, it is not known whether this reflects a long-term decline or just fluctuations related to the severity of winters in the respective periods (Laursen et al. 1997).

Despite being gregarious and preferring coastal waters the species is not particularly sensitive to hunting related disturbances. The present level of hunting is probably not influencing population size (Madsen et al. 1996)

Main threat is decreased habitat quality due to eutrophication reducing macrophyte cover.

Significance of the Danish population
163,900 equals 11% of total estimated northwest European winter population.

Population trend within Danish SPAs and SPA coverage
Most recent counts are lower than at designation time. Most of the observed changes are probably just weather-related fluctuations, but some (notably at SPA no. 43) are the result of decreased habitat quality as eutrophication has caused disappearance of/reduction in bottom macrophyte cover.

The coverage is high, as nearly all important wintering areas as shown in Jensen (1993) or Durinck et al. (1994) are included in the SPA network.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY Comments
1 42000 10500 1993  
24 15000 645 1993-95  
30 5000      
43 40000 3300 1993-94  
71 38000 15275 1994  
75 15000 6500 1993-94  
81 14000 17725 1987-89  
83 15000 8500 1995 Only part of area counted
84 10000 2190 1993-94  
85 25000 15300 1993-94  
86 7000      
88 19000 9800 1996  
89 25000 13900 1995  
96 11000 1645 1987-89  
105 25000 12499 1998  

 

Common Crane

Species name
Scientific: Grus grus
UK: Common Crane
DK: Trane

Red-list and SPEC status DK/Europe/World
DK: Rare
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
In Denmark, breeding occurs in blanket and raised bogs in heathlands and in wetlands in forest. Elsewhere a notable adaptation to man made landscapes has been noted during the last 30 years with breeding in small wetlands in fields and meadows (Tucker and Heath 1994). Breeding in such areas is still the exception in Denmark, though.

Occurrence in Denmark
The species is a breeding visitor and a passage visitor from other parts of Scandinavia.

Trends in Danish population - long-term and 1983-1998
The original Danish population went extinct around mid 19th century. The first definite breeding record in the 20th century was in 1952 (though breeding probably occurred at the site already in 1938 or 1936 (Løppenthin 1967)). Since then, an increase in the population has occurred to 6-10 pairs in 1993-96 and 11-12 pairs in 1997.

Main threat toward the Danish population is probably disturbance at nest sites.

Significance of the Danish population
The Danish population is very small compared to the European total of 52,793-77,970 pairs (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
The above table is not wholly up to date. The SPAs harbour most of the Danish pairs and the coverage probably is in the 80-90% range. The SPAs thus have a high importance for the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
6       0 1   1993-94  
7   0   1 1   1994  
13           40 1998  
17 P P   1 1   1993  
22       0 2   1993-94  
80       2 3   1995  
82           13 1995 Exceptionally 2000 staging 1998
89           35 1996  
Total 1 ?   4 8      

 

Eurasian Oystercatcher

Species name
Scientific: Haematopus oestragalus
UK: (Eurasian) Oystercatcher
DK: Strandskade

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Oystercatchers concentrates when foraging on sandy or stony intertidal flats.

Occurrence in Denmark
The species is a common breeder and an abundant passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
The total Westpalaearctic population has increased substantially during this century (Meltofte 1993) and the northwest European winter population is currently estimated at 874,000 birds (Rose and Scott 1997). The numbers in the by far most important Danish staging area – the Waddensea – fell during the 1980s because of decreased habitat quality caused by intensive mussel fisheries. Numbers have since at least partly recovered as a result of better regulation of the mussel fishery.

Significance of Danish population
More than 50,000 birds or nearly 6% of population total can occur simultaneously in Denmark.

Population trend within Danish SPAs and SPA coverage
Staging numbers within Danish SPAs have been mostly stable, though numbers fell in the Waddensea in a period of intensive mussel fishery in the 1980s. All major staging areas are within the SPA network. Coverage is thus high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
52 17000 15000 1993-94
53   16400 1992-94
55 5270 5385 1996
57 65000 39002 1987-89
60   6528 1992-94
65 13800 3900 1991-95

 

Avocet

Species name
Scientific: Recurvirostra avosetta
UK: Avocet
DK: Klyde

Red-list and SPEC status DK/Europe/World
DK: Yellow listed as AT
Europe: Secure (breeding), Localized (winter), SPEC 4 (breeding), SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Breeds at coastal sites with intertidal mud- or sandflats.

Occurrence in Denmark
The species is mainly a breeding visitor but also occurs as passage visitor from Sweden and Finland. In the moulting period also birds breeding in Northern Germany occur in the Danish part of the Waddensea (Meltofte 1993).

Trends in Danish population - long-term and 1983-1998
Until mid 19th century the species was a common breeding bird in Denmark. During the period 1860-1920 the population decreased dramatically mostly due to hunting. Since total protection in 1922, the population has increased from 700 pairs to around 5,000 pairs. During the 1990s local population decreases have occurred but the overall population increase has continued (Grell 1998).

Significance of the Danish population
Denmark holds an important part of the estimated European total of 27,762-38,436 pairs (Hagemeijer and Blair 1997). The 6,000-7,500 birds at moulting aggregations in the Danish part of the Waddensea (Meltofte 1993) are also of high importance.

Main threats at a European level are loss and disturbance of breeding habitat and the deterioration of feeding conditions at breeding and wintering sites (Tucker and Heath 1994).

Population trend within Danish SPAs and SPA coverage
The trend within the SPAs is almost identical to the overall trend for Denmark as the coverage is high - probably around 95% - and the SPAs are thus of the highest importance for the continued positive development in the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1 100 150   156 156   1993  
8 50 50   30 40   1992-94  
9 50 50            
10 170 170 600 285 285 1000 1987,
1987-91
 
12 8 8   10 10   1996  
13 15 17   172 172   1995  
15 20 20   110 110   1997  
20       1 1   1995  
23 200 200 1000 12 37 85 1996 Breeding numbers are from 1987-88
24 4 10 400     27 1995  
25 50 50   5 5   1995  
26 15 50   6 11   1987-89  
27 30 50   16 16   1996  
30       0 2   1988-89  
31 25 25   16 18   1993-95  
36 80 80   25 25   1996  
38 125 140   320 397   1993-96  
39 130 130 650 60 70 245 1998  
40 P P   6 6   1994  
41 2 2   0 0   1988-89  
43 335 335   592 670   1993-94  
47 50 50   7 7   1996  
49     300 2 2 2000 1991-94,
1993-94
 
51 P P   300 300   1993-94  
52       56 56   1997  
53 10 10   48 48   1997  
55 10 10 400 112 112 255 1997 Only counted in part of area 1997
57 650 750 7700 1066 1066 6500 1991,
1992-94
 
60 108 108   268 268 1313 1996 Staging numbers are from 1992-94
65 30 50   49 49   1996  
67       23 23   1997  
71 125 125   170 180   1996  
75 60 60   53 53   1994  
76 50 50   56 56   1996  
81 50 100   90 90   1995  
83 25 30   49 49   1993-96  
84       0 1   1985-89  
85 75 75   36 41   1996-98  
86       5 5   1985-89  
88 30 30   10 10   1997  
89 110 110   124 124   1994  
94 3 3   0 0   1993-94  
95       22 22   1988  
96 170 250   178 193   1987-89  
99           465 1996  
100       5 5   1993-94  
105 75 75   101 101   1997  
110 60 120   225 225   1997  
111 P P   4 5   1995  
Total 3100 3543   4881 5122      

 

Grey Plover

Species name
Scientific: Pluvialis squatarola
UK: Grey Plover
DK: Strandhjejle

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: (Secure), non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Grey Plovers forage on intertidal mud- and sandflats, young birds also to some extent on short grassed meadows (Grell 1998).

Occurrence in Denmark
The species occurs as passage visitor and also in low numbers as winter and non-breeding summer visitor.

Trends for relevant population(s) overall and in Denmark
There is no information regarding trends in the East Atlantic winter population, the size is currently estimated at 168,000 birds (Rose and Scott 1997). The trend in Denmark is likewise unknown.

Significance of the Danish population
Internationally important numbers in Denmark only occur at few high tide roosts in the Waddensea. Up to 5,000 birds (Asbirk et al. 1997) or 3% of population total can occur simultaneously in Denmark.

Population trend within Danish SPAs and SPA coverage
Trends within Danish SPAs are not known, but numbers are probably reasonably stable. The two SPAs above are probably the only Danish sites that harbour internationally important concentrations.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
53   2872 1992-94
57 3000 5550 1992-94

(Common / Northern) Lapwing

Species name
Scientific: Vanellus vanellus
UK: (Common/Northern) Lapwing
DK: Vibe

Red-list and SPEC status DK/Europe/World
DK: Yellow list in category X (breeding)
Europe: (Secure), non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Lapwings occur in a variety of open habitats such as inter tidal flats, saltmarshes, pastures and agricultural fields.

Occurrence in Denmark
The species is a breeding and passage visitor, some winter in mild winters.

Trends for relevant population(s) overall and in Denmark
Breeding populations have decreased recently in several European countries (Snow and Perrins 1998) probably due to changes in land use, but overall trend for European population – estimated at 7,000,000 birds (Rose and Scott 1997) - is not clear. Breeding numbers in Denmark are decreasing (Grell 1998) while the trend for staging birds is unclear (Asbirk et al. 1997).

Staging birds will probably be benefited by the creation of the many new hunting and disturbance free core zones as the species responded positively to the experimental reserves (Madsen et al. 1995).

Significance of the Danish population
The 200,000 birds regularly staging in Denmark in autumn equals nearly 3% of European total.

Population trend within Danish SPAs and SPA coverage
There is no further information regarding trends in staging numbers in Danish SPAs. Most major sites mapped in Meltofte (1993) are within the SPA network, though a few important ones such as Sjørring Sø are outside. The coverage is thus high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
60 39500 13000 1992-94

 

(Red) Knot

Species name
Scientific: Calidris canutus
UK: (Red) Knot
DK: Islandsk ryle

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Localized, SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Knots occur mostly in large, dense flocks at extensive tidal flats, often predominantly of muddy sands (Tucker and Heath 1994).

Occurrence in Denmark
The species occur as passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
Two different populations and subspecies occur in Denmark. The ssp. islandica breeding in Canada and Greenland winter mainly in northwest Europe, while Siberian birds of the nominate subspecies pass through on their way to winter quarters in western and southern Africa.

The northwest European winter population dropped during the 1970s probably because of unfavourable conditions in breeding areas. Despite partial recovery, numbers in early 1990s remain around 40% lower than in early 1970s at 310,000 birds. In the same period a major redistribution of wintering birds from the Waddensea to estuaries in the U.K. took place probably as a result of decreased food supply in the Waddensea following intensification of mussel fisheries (Tucker and Heath 1994). Rose and Scott (1997) gives the population as 345,000 birds while Snow and Perrins (1998) reports that recent information from U.K. and the Waddensea suggests that population is around 500,000. The Danish part of the Waddensea holds up to 50,000 birds from this population, highest numbers occurring in spring. High numbers have only been reported from 1970s and onwards and probably did not occur prior to that time (Meltofte 1993). The present trend in Danish staging numbers is unknown (Asbirk et al. 1997)

The Siberian population of 516,000 birds (Rose and Scott 1997) mainly stage in the German part of the Waddensea but up to at least 15,000 birds can occur in the Danish part (Meltofte 1993).

Main threats toward the population(s) on European level are habitat loss through land claims, reduction in food supply due to intensive mussel fisheries and disturbance from overflying aircraft and from recreational activities (Tucker and Heath 1994).

Significance of the Danish population
50,000 birds equal 10-16 of total population of ssp. islandica.

Population trend within Danish SPAs and SPA coverage
There is probably little change in the numbers staging within the SPAs.

All major staging sites are SPAs. The coverage is hence high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
52   20000 1993-94
53   7575 1992-94
57 5000 24170 1992-94
60   21310 1992-94

 

Sanderling

Species name
Scientific: Calidris alba
UK: Sanderling
DK: Sandløber

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: None

General ecology and habitat requirements
Staging Sanderlings forage along sandy beaches and on inter tidal flats.

Occurrence in Denmark
The species occurs as passage and winter visitor.

Trends for relevant population(s) overall and in Denmark
Trends are not known, but the total East Atlantic/West and South African winter population is estimated at 123,000 birds (Rose and Scott 1997). This "population" involves both Siberian and Greenland breeders (Meltofte 1993). Up to at least 4,500 occur simultaneously in Denmark; the national trend is unknown (Asbirk et al. 1997).

Significance of the Danish population
4,500 birds equal 3-4% of population total.

Population trend within Danish SPAs and SPA coverage
Trends within SPAs are unknown; there are large local fluctuations related to weather and the amount of beached seaweed especially in winter (Meltofte 1993).

The coverage is high. Internationally important numbers only occur in the Waddensea region, which is well covered though one important roost - at Blåvandshuk at the extreme northern edge of the area – is outside the SPA network.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
53   2199 1992-94
57   3000 1992-94

 

Dunlin

Species name
Scientific: Calidris alpina
UK: Dunlin
DK: Almindelig ryle

Red-list and SPEC status DK/Europe/World
DK: Rare (the breeders of ssp. schintzii) and Yellow listed as AY (ssp. schintzii) and AT (both subspecies)
Europe: Vulnerable, SPEC 3
World: Lower Risk (least concern)
Annexes: None

General ecology and habitat requirements
Staging Dunlins can occur in a variety of habitats, but numbers are highly concentrated at the very best (most productive) inter tidal flats (Meltofte 1993).

Occurrence in Denmark
Ssp. alpina is an abundant passage visitor (smaller numbers winter) and ssp. schintzii is an uncommon and decreasing breeder and a common passage visitor.

Trends for relevant population(s) overall and in Denmark
Wintering numbers in western Europe fell markedly 1970-90, despite partial recovery numbers in early 1990s are still below those of early 1970s (Tucker and Heath 1994).

Total European winter population has been given as at least 1,000,000 birds (Tucker and Heath 1994) and total population of ssp. alpina on the Siberia/Europe/West Africa flyway is estimated at 1,373,000 birds (Rose and Scott 1997). The Baltic/UK/Ireland population of ssp. schintzii numbers 21,000 birds (Rose and Scott 1997).

Trends in Danish staging numbers are unknown. Up 450,000 birds occur simultaneously during passage and 56,600 in winter (Asbirk et al. 1997).

The number of breeding birds in Denmark (ssp. schintzii) has decreased from an estimated 50,000-100,000 pairs 100 years ago to presently approximately 450 pairs. Numbers are still decreasing except where specialised management is applied (Grell 1998).

Threats against habitats for staging birds mentioned on a European scale are habitat destruction as a result of land-claim and spread of the invasive grass Spartina and large scale pollution incidents. Breeding numbers of ssp. schintzii have been reduced as a result of land-use changes, agricultural intensification or abandonment and afforestation of moorland (Tucker and Heath 1994).

Significance of the Danish population
Up to 33% of the total ssp. alpina population on the Siberia/Europe/West Africa flyway can occur simultaneously in Denmark.

Population trend within Danish SPAs and SPA coverage
Numbers staging within SPAs seem mostly stable. All important staging areas mapped in Meltofte (1993) are SPAs. Coverage is thus high.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1 1 1 15000     4420 1993  
2           24000 1993-94  
9     560          
10     45000     40000 1994  
38 35 40 15000     12440 1996 Only part of area counted 1996
39 35 65            
43 70 70   200 250   1993-94  
47 5 10            
52 1 1 72500     75000 1993-94  
53     2025000     88500 1992-94  
55 6 6 25550     14800 1996  
57 30 40 360000     364929 1992-94  
60           62000 1992-94  
65 20 25 32500         No new data

 

Ruff

Species name
Scientific: Philomachus pugnax
UK: Ruff
DK: Brushane

Red-list and SPEC status DK/Europe/World
DK: Vulnerable
Europe: Secure, SPEC 4
World: Lower Risk (least concern)
Annexes: I, II/2

General ecology and habitat requirements
In Denmark, the species breed at short-grazed meadows often with brackish water.

Occurrence in Denmark
The species is a breeding visitor and a more numerous passage visitor from the rest of Scandinavia and to a lesser extent Siberia (Meltofte 1993).

Trends in Danish population - long-term and 1983-1998
Main threat toward the population is habitat changes, which include both agricultural intensification and abandonment, as the species seems to have a very narrow habitat preference. Possibly hunting during migration and in wintering areas is also of some significance.

The Danish population has decreased continuously probably since the beginning of the 20th century though population estimates from the beginning of the century do not exist (Grell 1998). The species has disappeared from almost all inland sites and now almost exclusively breeds in saltmarshes and other coastal localities.

The population was estimated at 840-1,104 "pairs" during the period 1960-69, at 500 females in 1980 and at 750 breeding females in the late 1980s. The increase during the 1980s and the possibly almost stable situation at present are exclusively the result of specialised management at the key site Tipperne (SPA no. 43). The population here increased from 120 breeding females in 1980 to about 350 breeding females in the late 1980s. Most recently, though, the numbers have decreased again and were much lower 1995-97 (Laursen 1999). At most other sites, the decrease continues (Grell 1998). The total Danish population was estimated at 500 breeding females by Grell (1998), but if numbers at the key site Tipperne (SPA no. 43) do not recover from the low numbers of 1995-97, the population might be considerably smaller.

Significance of the Danish population
In pure numbers, the Danish population is only of minor importance. Eventual disappearance of this and other declining populations in the southern part of the species range will, though, signify a continuation of the already marked range contraction (Tucker and Heath 1994). Total European breeding population is estimated at 1,105,655-10,139,209 of which 1,000,000-10,000,000 million are in Russia (Hagemeijer and Blair 1997).

In Denmark passage visitors occur in much higher numbers than the breeders do. As total flyway population is believed to be very large, though, no Danish site satisfies the 1% criterion though aggregations of up to 14,000 have been recorded occasionally (Meltofte 1993).

Population trend within Danish SPAs and SPA coverage
Apart from the positive situation at SPA no. 43 (though large year-to-year fluctuations occur) the species has decreased at almost all sites and disappeared as a breeding bird from several. The situation outside SPAs is even worse and the coverage is now probably near 100%. The management of SPAs is thus of crucial importance for the future of the Danish breeding population. The main aggregations of passage visitors also occur within SPAs.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1 25 25   11 11   1993  
7       0 0   1993  
8       0 0   1992-94  
10       13 13   1987  
13       36 36   1995  
15       0 3   1993  
20       4 4   1995  
21       0 2   1987-89  
23 45 45   16 44   1987-88  
24 1 4           Probably no longer breeds
25 P P   0 0   1988  
27 P P   1 1   1994  
38 10 25   2 2   1994  
39 P P   P? P?   1998  
41 1 1   0 4   1988-89  
43 150 150   100 150   1993-94  
49 15 15   0 0   1996  
51       5 10   1993-94  
52 12 12         1991 Possibly almost gone
53 P P   0 0   1997  
55 4 4   0 4   1987-89  
57       26 26   1991  
60       5 5 3815 1996  
61 P P   0 0   1993-94  
65 40 50   4 4   1997  
67 10 11   0 0   1997  
71       0 2   1990-96  
76 P P   0 0   1994  
82 12 15   0 0   1993  
85 1 1   0 0   1998 Last breeding 1988-89
89 10 10   2 2   1993-96  
91       0 0   1993-94  
95       0 22   1988  
99 P P           No new data, probably no longer breeds
110       40 40   1995  
111       0 0   1995 Last possible breeding 1993
Total 331 363   277 397      

 

Bar-tailed Godwit

Species name
Scientific: Limosa lapponica
UK: Bar-tailed Godwit
DK: Lille kobbersneppe

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Localized, SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: I, II/2

General ecology and habitat requirements
Staging Bar-tailed Godwits prefers sheltered bays and estuaries with sand or muddy inter tidal flats.

Occurrence in Denmark
The species is a passage visitor and in lower numbers a winter visitor.

Trends for relevant population(s) overall and in Denmark
Two populations are involved both belonging to the nominate subspecies. Birds from northern Scandinavia and western parts of Russia winter in Europe, while birds from further east pass through on their way to wintering grounds in Africa.

The Westpalaearctic winter population fluctuated 1970-90 (Tucker and Heath). This population numbers 115,000 birds while the birds on the Siberian/west African flyway number 700,000 (Scott and Rose 1997).

Up 75,000 to birds occur simultaneously in Denmark during peak of spring passage, trends are unknown (Asbirk et al. 1997).

Main threats towards the population(s) at the European level are land-claim, pollution reducing habitat quality and human disturbance (Tucker and Heath 1994).

Significance of the Danish population
The 75,000 birds occurring simultaneously equal more than 9% of the combined total for the involved populations.

Population trend within Danish SPAs and SPA coverage
Numbers staging within Danish SPAs seem to be reasonably stable. All major staging areas as mapped in Meltofte (1993) are SPAs. Coverage is hence high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
23 3800 63 1996
36 3000 4630 1995
38 6500 2050 1996
43 3500 1000 1993-94
49 400    
52 30000 25000 1993-94
53 5000 16100 1992-94
55 3000    
57 56000 62084 1992-94
65 5400    

 

Curlew

Species name
Scientific: Numenius arquata
UK: Curlew
DK: Stor regnspove

Red-list and SPEC status DK/Europe/World
DK: Rare (breeding)
Europe: Declining, SPEC 3 (winter)
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Curlews occur mainly on intertidal flats and in saltmarshes. In lower numbers also in other, open habitats. The small Danish breeding population occur mainly in heathlands and saltmarshes.

Occurrence in Denmark
The species is a scarce breeding visitor and a more numerous passage visitor. A few thousand birds winter, mainly in the Wadden Sea (Meltofte 1993).

Trends for relevant population(s) overall and in Denmark
The total European population (equivalent to the entire population of ssp. arquata) is estimated at 348,000 birds (Rose and Scott 1997). The population is declining (Tucker and Heath 1994, Hagemeijer and Blair 1997).

The decline in the European population is probably mainly a result of habitat loss or degradation in breeding areas because of agricultural intensification and afforestation. During migration, the species is threatened by building development and disturbance at high-tide roosts, locally by chemical pollution and hunting (Tucker and Heath 1994). Intensive mussel fisheries can also be detrimental to the species as it often feeds on polychaete worms associated with the mussel banks.

Staging Curlews are very vulnerable to disturbance (Grell 1998, Christensen et al. 1999)

Significance of the Danish population
Normally the total number of birds simultaneously staging in Denmark does not exceed 10,000 (Meltofte 1993). In the most important Danish staging area, the Wadden Sea, numbers of staging Numenius spp. have increased 1990-95, probably as result of lower pressure from hunting and other disturbances (Laursen and Frikke 1997).

Population trend within Danish SPAs and SPA coverage
Only in the Wadden Sea (SPA no. 57) do concentrations of international importance occur in Denmark. Coverage is thus high.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY
6 6 8          
17 2 3          
18 2 2          
19 2 2          
22 5 10          
37 10 30   14 18   1996
43 6 6          
57     5600     3325 1992-94
65 25 25   26 26   1996
91 1 2          
110 5 15   68 68   1996
111 P P          

 

Spotted Redshank

Species name
Scientific: Tringa erythropus
UK: Spotted Redshank
DK: Sortklire

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Spotted Redshanks occur at sheltered sites with soft substrates by the coast and in lower numbers also at inland sites.

Occurrence in Denmark
The species is a passage visitor to Denmark.

Trends for relevant population(s) overall and in Denmark
The birds occurring in Denmark breed in Norway, Sweden, Finland and northwest Russia and winter mostly in west Africa, though some stay in west Europe and the Mediterranean. No overall trend is known for this population, though the substantial Finnish part is believed to have decreased (Snow and Perrins 1998). The population total is estimated at 75,000-150,000 (Rose and Scott 19979.

Significance of the Danish population
The majority of the birds pass through Denmark quickly and internationally important numbers therefore rarely occur. A few more sites might qualify if turnover rates were known. Up to 2,500 birds equalling 2–3% of population total can stage simultaneously in Denmark (Asbirk et al. 1997).

Population trend within Danish SPAs and SPA coverage
Trends within SPAs are not known. Only in the artificial lagoon "Saltvandssøen" in SPA no. 60 numbers do totals reach internationally important levels. Also most sites of secondary importance are SPAs and coverage is thus high.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
60   1269 1992-94

 

Redshank

Species name
Scientific: Tringa totanus
UK: Redshank
DK: Rødben

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Declining, SPEC 2
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Redshanks mainly occur at muddy inter tidal flats. The birds of the Icelandic ssp. robusta, though, prefer more sandy or stony sites (Meltofte 1993).

Occurrence in Denmark
The species is a breeding visitor, passage visitor and (mainly Icelandic. ssp. robusta) winter visitor.

Trends for relevant population(s) overall and in Denmark
The breeding population of the nominate subspecies has declined in several European countries. Midwinter counts, though, show stable numbers (Tucker and Heath 1994). The total number of ssp. totanus on the Eastern Atlantic flyway is estimated at 177,000 (Rose and Scott 1997). Southern breeders of this population mainly winter in the west Mediterranean though some remain in northwest Europe while most of the northern breeders winter in west Africa (Meltofte 1993). Icelandic birds – numbering 109,000 birds (Rose and Scott 1997) - winter in northwest Europe mainly in the UK.

Up to 15,000 birds occur simultaneously during passage and up to 2,000 winter, trends in staging numbers are unknown (Asbirk et al. 1997). Breeding numbers have declined almost constantly this century, with the exception of a few sites where specialised management has been shown effective in reversing trends. The breeding population is believed to be 10,000-15,000 pairs (Grell 1998).

Threats affecting staging areas are habitat destruction and deterioration as a result of land-claim, industrial development, dredging, pollution, human disturbance and spread of Spartina on inter tidal flats (Tucker and Heath 1994).

Significance of Danish population
15,000 birds equal 5% of the total for both involved populations. Undoubtedly a much higher percentage utilises Danish areas within a year.

Population trend within Danish SPAs, SPA coverage and significance of SPAs
The material is too small to judge possible trends in staging numbers within SPAs. The coverage and significance are both high though as all major sites mapped in Meltofte (1993) are SPAs.

Occurrence in Danish SPAs

Area no. 1983 NSB Now NSB NDY
57 10300 1680 1992-94
65 2200 175 1991-95

 

Greenshank

Species name
Scientific: Tringa nebularia
UK: Greenshank
DK: Hvidklire

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: II/2

General ecology and habitat requirements
Staging Greenshanks mainly occur at mud-, sandflats, and coastal lagoons and in smaller numbers also at inland sites.

Occurrence in Denmark
The species is a passage visitor to Denmark.

Trends for relevant population(s) overall and in Denmark
The European population apparently fluctuates and overall trend is not clear (Snow and Perrins 1998). The total size of the population on the Europe/West Africa flyway is only known within broad limits: 100,000-1,000,000 birds (Rose and Scott 1997). The total number of birds staging simultaneously is up to 6,400 (Asbirk et al. 1997).

Significance of the Danish population
The only count of numbers of international significance was in the Waddensea (SPA no. 57) in 1990 with 3,400 individuals (Meltofte 1993). As the birds pass through quickly the number of birds utilizing a given site during a season is significantly higher than the max. counts.

Population trend within Danish SPAs and SPA coverage
As birds usually only stage for a short time there are large random fluctuations in numbers counted. Most important staging sites are SPAs and the coverage is thus high.

 

Wood Sandpiper

Species name
Scientific: Tringa glareola
UK: Wood Sandpiper
DK: Tinksmed

Red-list and SPEC status DK/Europe/World
DK: Vulnerable
Europe: Declining, SPEC 3
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
The species breeds in bogs and other wetlands in heathlands preferably far from trees.

Occurrence in Denmark
In Denmark the species is breeding visitor and a more numerous passage visitor from the large populations in the rest of Scandinavia (Meltofte 1993).

Trends in Danish population - long-term and 1983-1998
Until the beginning of the 20th century the species was a reasonably common and widespread breeder in heathlands in Jutland (Grell 1998), while the species has most probably never been a regular breeder in the Danish Isles (Løppenthin 1967).

Following large scale conversion of heathlands and bogs to other types of landscapes the species decreased rapidly and the first detailed inventory of the species, which took place during the 1950s, yielded an estimate of 200-300 pairs at 44 sites. Later inventories recorded 100-150 pairs at 59 sites (1960-69), 88-112 pairs at 53 sites (1978-81) and 53-64 pairs at only 20 sites (1982-84). The 1982-84 survey furthermore showed that there were large year-to-year fluctuations in breeding numbers. Only 9 sites were occupied in all three years (Grell 1998).

The population during the years 1993-96 was estimated at 63-82 pairs at 12-13 sites. The population, thus, seems to have stabilised or maybe even recovered slightly, though the apparently positive tendencies might only be a result of better coverage of the key sites in Northwest Jutland (Grell 1998).

The main threat at a European level is destruction of breeding habitat through drainage followed by afforestation or conversion to agriculture and extraction of peat for fuel or for soil improvement in gardens (Tucker and Heath 1994). In Denmark, disappearance from some former breeding sites has been attributed to disturbance caused by recreational activities (Jensen 1995).

Significance of the Danish population
The Danish population is very small in comparison with the European total of 398,842-1,412,474 pairs (Hagemeijer and Blair 1997). Eventual disappearance of the Danish and the other small remnant, populations in the South part of the species breeding range will nevertheless mean a marked range contraction.

Most passage visitors occur in the Danish isles (Meltofte (1993), but due to the total population being very large and the fact that that most visitors only stay for shorter periods, no Danish locality ever approaches internationally important numbers.

Population trend within Danish SPAs and SPA coverage
Probably the apparent increase in numbers in Danish SPAs since designation is mostly an artefact due to better data quality in recent years. There might be some real increase in a few core areas, though, but the species has disappeared from a number of smaller localities. The SPA coverage is high, probably above 90%, and the management of SPAs, especially the few key sites harbouring the majority of the population, are thus of crucial importance for the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NDY
NSB
Comments
5 0 2   0 1   1993-94
6 2 5   0 0   1993-94
7 3 4   0 0   1993
10 1 2   0 0   1987
17 P P   13 15   1993-94
18 3 >3   0 0   1993-94
22 8 12   35 44   1993-94
37 5 10   7 7   1995
42 4 4   0 0   1995
44 2 2   0 0 110 1995
46 1 2   0 0   1993
48 2 3   0 0   1995
No breeding since 1980s
50 3 5   1 2   1993-94
54 0 2   0 0   1995
Last breeding around 1970
56 1 2   0 0   1993-94
57 1 3   0 0   1991
58 1 1   0 0   1989
61 5 5   3 6   1996
66       1 2   1994
69 0 3   0 0   1993-94
70 2 4   0 0   1993-94
91       0 0   1993-94
Total 45 72   60 77    

 

Gull-billed Tern

Species name
Scientific: Gelochelidon nilotica
UK: Gull-billed Tern
DK: Sandterne

Red-list and SPEC status DK/Europe/World
DK: Endangered
Europe: (Endangered), SPEC 3
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
In Denmark the species breeds on small, sparsely vegetated islands and islets near suitable foraging areas such as tidal flats or saltmarshes (Grell 1998). Grell (1998) also states that formerly also large, open dry heathlands were important foraging areas, but this is no longer the case. The species still utilise the coastal heath - dune area south of Lakolk (SPA no. 66) for foraging, though.

Occurrence in Denmark
The species is a breeding visitor in Denmark.

Trends in Danish population - long-term and 1983-1998
Main threats toward the species in Europe include loss and degradation of foraging habitats through agricultural intensification including drainage and use of pesticides both decreasing prey levels and directly killing birds. Inappropriate water level management allowing easier access for various mammalian predators or flooding, egg-collecting for consumption and human disturbance are other detrimental factors (Tucker and Heath 1994).

From mid 19th century until around 1950 the Danish population fluctuated between 150 and 500 pairs, since then an almost constant population decline and range contraction have occurred to only 13-16 pairs in 1993-96 (Grell 1998). The breeding success has in recent years been very low and the species probably only survives as a Danish breeder because of immigration from the more successful German population (Rasmussen and Fischer 1997).

Significance of the Danish population
The Danish population is only a very small part of the European total of 4,922-8,551 (Hagemeijer and Blair 1997). Eventual loss of this and the likewise small (50-60 pairs, Tucker and Heath 1994) German population will, though, mean a marked contraction of the species European range.

Population trend within Danish SPAs and SPA coverage
Until the mid-1980s, the Danish colonies were situated more northerly in Jutland. Deteriorated habitat quality at some important foraging areas and (in SPA no.38) immigration of Fox (Vulpes vulpes) to an important breeding island at that time caused a shift of breeding sites to the Waddensea area (Grell 1998). The coverage is 100%. Probably the total population also bred in SPAs before the shift to the Waddensea region. The management of the few SPAs still holding this species is of paramount importance for the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
52       1 1   1997  
55       7 7   1997  
65       3 3   1997 No breeding success 1997
Total       11 11      

 

Sandwich Tern

Species name
Scientific: Sterna sandvicensis
UK: Sandwich Tern
DK: Splitterne

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Declining, SPEC 2
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
Breeds on small sparsely vegetated island and islets mainly by the coast and almost always in conjunction with colonies of Black-headed Gull (Larus ridibundus).

Occurrence in Denmark
The species is a breeding visitor and in smaller numbers a passage visitor from the small populations in the rest of Scandinavia. Probably birds from Northern Germany are also part of the flocks of moulting birds of up 5,000 occurring at Blåvandshuk (SPA no. 50) (Grell 1998).

Trends in Danish population - long-term and 1983-1998
The country probably held a population of 7,000-8,000 pairs until around 1950. A marked decline occurred from then until the early 1970s, where the population was approximately halved. The reduction was caused mainly by egg collecting for consumption and other disturbance, the increase in the Herring Gull (Larus argentatus) population. Possibly decreased prey availability as result of intensive fishery on sandeels (Ammodytidae) was also partly responsible for the observed decline (Ferdinand 1980). After some recovery in the 1980s, a decrease in the main colony at Hirsholmene (SPA no. 11) occurred for reasons unknown and in 1993-96, the total Danish population was estimated at 4,500 pairs (Grell 1998). Main threats toward the Danish population are disturbance, invasion of predators including the alien Brown Rat (Rattus norvegicus) and American Mink (Mustela vison) on formerly predator-free islands and possibly the decrease in the Danish population of Black-headed Gull (Grell 1998).

Significance of the Danish population
The Danish population amounts to 3-4% of the European total of 119,561-141,784 pairs (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
Overall the population within SPAs has been almost stable, but much change in numbers has occurred within the specific areas. For instance the important colony at Hirsholmene (SPA no. 11) held 1400 pairs at designation, 3,000 pairs in 1988 and 1356 pairs in 1994. Around 1940 the colony held no less than 4,000 pairs. The other main colony at Langli in SPA no. 55 has increased rapidly in recent years (Laursen 1999) and held as many as 1,600-1,800 pairs in 1999 (Thor Bue Hansen pers. com).

The coverage is high, nearly 95%, and proper management of SPAs, especially regarding protection from human disturbance is therefore of utmost importance for the Danish population of the species.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY/ Comments
1 20 20   22 22   1993
2 200 200   0 1   1994
9 100 100   0 0   1993
10       60 60   1987
11 1400 1400   1356 1356   1994
23 150 300   0 0   1987-88
31           0 1995
36 300 300   0 0   1994
38 1650 1650   1050 1520   1994-96
55 0 0   1355 1355   1997
57       0 0   1991
65 P P   0 0   1997
No breeding after 1987
71 100 100   100 140   1996
75 100 200   300 300   1994
76       78 78   1994
81       1 1   1995
83 50 200   0 0   1993-96
84       200 200   1985-89
88       40 40   1997
89 200 200   170 170   1993-96
96 P P   0 0   1988
98 400 400   16 16   ?
110       1 1   1993 Probably no longer breeds
  4672 5072   4749 5260    

 

Common Tern

Species name
Scientific: Sterna hirundo
UK: Common Tern
DK: Fjordterne

Red-list and SPEC status DK/Europe/World
DK: Not listed
Europe: Secure, non-SPEC
World: Lower Risk (least concern)
Annexes: I

General ecology and habitat requirements
The species breeds both at coastal localities, on islands and islets and by freshwater lakes.

Occurrence in Denmark
The species is a breeding visitor and an abundant passage visitor from the large populations in Norway, Sweden and Finland.

Trends in Danish population - long-term and 1983-1998
The trend in the Danish population before the 1960s is not well known as many observers were unable to tell the present species from Arctic Tern (Sterna paradisaea), but the population is believed to have decreased since the 1940s. The population was estimated at 600-800 pairs in the period 1970-76, at 1,500 pairs in 1988 and at 1,000 pairs in 1993-96. The species fluctuates markedly from year to year, but the increase (recovery) during the 1980s was apparently the result of the creation of numerous island reserves with entry prohibited during the breeding season. The reasons for the renewed decrease are unclear, possibly there are some longer term fluctuations involved (Grell 1998).

Probably the main threat toward the Danish population still is disturbance at breeding sites. Too high vegetation on breeding sites as result of abandonment and reduced food availability as a result of over-exploitation of fish stocks and eutrophication of lakes and coastal waters might also play some role.

Significance of the Danish population
The Danish breeding population is only a small part of the European total of 225,105-287,250 (Hagemeijer and Blair 1997).

Population trend within Danish SPAs and SPA coverage
The apparent increase in numbers within SPAs is probably partly a result of better coverage. Several colonies were at designation just registered as P for present in the new data these colonies appear with exact numbers. Some larger colonies are known to have become established or to have increased during the period, though. Some local disappearances or decreases have also been noted. The population within SPAs is probably much the same or slightly higher than at designation.

Though many breeding sites are outside SPAs (Grell 1998), these typically harbour single pairs or small colonies and the coverage is probably not below 90%, and proper management of SPAs is therefore of crucial importance for the Danish population.

Occurrence in Danish SPAs

Area no. Min. BP 1983 Max. BP NSB Min. BP Now Max. BP NSB NDY Comments
1 0 0   45 45   1993  
9 20 20   16 16   1993  
10       5 6   1987  
11 25 50   175 175   1994  
13       1 1   1994  
15 20 20   10 10   1997  
20       20 20   1995  
21       1 3   1987-89  
23 40 40   0 0   1987-88  
30       0 6   1988-89  
38 50 50   140 140   1996  
39 P P   12 12   1998  
43 11 18   12 12   1993-94  
47 15 15   0 0   1988  
52       72 72   1997  
55 P P   4 4 18,000 1997